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Llerasia Triana, Ann. Sci. Nat. Bot. 4, 9: 37 (1858).

Chrysothamnus Nutt. sect. Diplostephioides Benth. & Hook. f., Gen. Pl. 2(1): 255 (1873). Type: not cited. Lectotype (selected by Bake, 1927: 108): Haplopappus hypoleucus Turcz. = Llerasia hypoleuca (Turcz.) Cuatrec.

Bigelowia DC. sect. Diplostephioides (Benth. & Hook.f.) A. Gray, Proc. Amer. Acad. Arts 8: 638 (1873).

Neosyris Greene, Erythea 3: 115 (1895). Type: not stated.

Haplopappus Cass. sect. Diplostephioides (Benth. & Hook.f.) S. F. Blake, Amer. J. Bot. 14(3): 108 (1927).

Lectotype (selected by Cuatrecasas, 1970: 40): Llerasia lindenii Triana


Blake, S. F. (1927). The Section Diplostephioides of Aplopappus. Amer. J. Bot. 14(3): 107115.

Cuatrecasas, J. (1970). Reinstatement of the genus Llerasia (Compositae). Biotropica 2: 3945.

Cabrera, A. L. (1982). Dos Compuestas nuevas de Bolivia. Hickenia 1(53): 281284.

Hall, H. M. (1928). The genus Haplopappus: A phylogenetic study in the Compositae. Carnegie Inst. Wash. Publ. No. 389. [Sect. 21 Diplostephioides pp. 355360].

Key to species [modified from Cabrera (1982)]

1.Capitula with 713 florets L. soratensis

Capitula with 35(6) florets 2

2. (1) Leaves and branches completely glabrous L. boliviensis

Leaves densely woolly or lanuginous beneath; branchlets woolly, lanuginous or glabrous 3

3. (2) Florets 5; leaf lamina 34.7 0.81.4 cm, elliptic or oblong-elliptic, short-attenuate, apices acute to obtuse; venation inconspicuous beneath L. ledifolia

Florets 34; leaf lamina lanceolate or elliptic lanceolate, long-attenuate, apices acute; secondary veins conspicuous beneath or hidden by indumentum4

4. (3) Leaves large, 10.512.5 2.83.5 cm; secondary veins conspicuous beneath L. lucidula

Leaves much smaller, 56 0.71 cm; secondary veins hidden by indumentum beneath L. beckii

Llerasia beckii Cabrera, Hickenia, 1(53): 281 (1982). Type: Bolivia. Depto. La Paz, Prov. Nor Yungas: camino de La Paz a Coroico a 72 km de La Paz, 3.050-3.150 m s.m. Ceja de la montaa, bosque cori ceo en zona muy nebulosa. Leg. St. G. Beck No. 1846, 28-VII-1979. Holotype: SI; isotypes: LPB, M.

Bolivia (La Paz).

Llerasia boliviensis (Cabrera) Cuatrec.,Biotropica, 2: 43 (1970)

Haplopappus (Diplostephioides) boliviensis Cabrera, Blumea 7(1): 193 (1952). Type: [Bolivia:] Hab.: Liane an der Waldgrenze ber Tablas, 3400 m alt., Mai 1911, [Herzog] n. 2186, Bl. orange, nach Hyacinthen duftend (Typus). Holotype: L; isotypes: LP (898904 according to Freire & Iharlegui, 2000: 321, but see Obs.), S.

Bolivia (Cochabamba).

Llerasia ledifolia (S. F. Blake) Cuatrec., Biotropica 2: 45 (1970).

Aplopappus ledifolius S. F. Blake, Amer. J. Bot. 14(3): 112 (1927). Type: .'Bolivia (?): Huaycani, alt. 3050-3355 m., May 1866, R. Pearce. Holotype: K; isotype: US (a fragment of the K specimen).

Bolivia (Cochabamba, La Paz), ?Peru.

Llerasia lucidula (S. F. Blake) Cuatrec., Biotropica 2: 43 (1970).

Moquinia macrocephala Rusby, Descr. New Sp. S. Amer. Pl. : 162 (1920). Type: Unduavi, North Yungas, Bolivia, 3,000 M., November, 1910. A shrub 3 or 4 M. high. (Buchtien, No. 3080.) Holotype: US.

Aplopappus lucidulus S. F. Blake, Amer. J. Bot. 14(3): 114 (1927). Type: Bolivia: Unduavi, North Yungas, alt. 3300 m., Nov. 1910, Buchtien 3080. Type in the U.S. Nat. Herb., nos. 1, 179, 267. [Nordyungas: Unduavi, 3300 m, Kletterstrauch 34 m, XI-1910, Buchtien 3080 according to Cuatrecasas, 1970: 43] This citation, by Blake, of the numbers is actually in error for 1179267, which is the holotype. Holotype: US (1179267).

*Gochnatia macrocephala (Rusby) Cabrera, Notas Mus. La Plata, Bot. 15(No. 74): 41 (1950).

Bolivia (La Paz).

Note: Cabrera (1971: 157) suggested that, following pollen analysis by Wodehouse (1929), the Buchtien collection showed closer affinity to Moquinia, suggesting M. racemosa (Spreng.) DC. [VERNONIEAE], a Brazilian species. This overlooked the fact that Blake had, in the interim, described the Buchtien collection as Aplopappus lucidulus, a taxon now accepted as belonging to Llerasia. Cuatrecasas (1970: 43) stated that the species was only known from the type, which is presumably where Rusby had described his Moquinia.

Bolivia (La Paz).

Llerasia soratensis (S. F. Blake) Cuatrec., Biotropica 2: 43 (1970).

Linosyris (Dolichogyne) mandonii Sch.Bip., Bull. Soc. Bot. France 12: 81 (1865), nom. nud. (based on Mandon 233)

Aplopappus soratensis S. F. Blake, Amer. J. Bot. 14(3): 109 (1927). Type: Bolivia: In rocky places in the subalpine and apline region near Sorata, between Nasacara and Lacatia, alt. 3400-3700 m., Aug.Nov, 1860, Mandon 233. Holotype: GH; isotypes: K, US (fragment of K).

Bolivia (La Paz).

Note: Bigelowia mandonii Benth. & Hook.f., Gen. Pl. 2(1): 255 (1873) although cited by Hall (1928: 357) and others, no such combination was proposed in Genera Plantarum providing the specific epithet combined with the generic name. The first time the combination is found is in Index Kewensis but is still based upon Schultz Bipontinus nom. nud. (via reference to Genera Plantarum), and can only be attributed to B. D. Jackson (q.v.). In Genera Plantarum the name actually appears in a list of species of Linosyris as Linosyris mandonii Sch.Bip., which might possibly to be considered as a new name, provided by Bentham & Hooker, for Haplopappus hypoleucus Turcz. The diagnostic characters provided by Bentham & Hooker are those distinguishing sect. Diplostephioides, and nothing more. Linosyris mandonii Sch.Bip. also appeared in Linnaea 34(5): 534 (Feb. 1866) as a nom. nud. Hall (1928: 357) was of the opinion that the appearance of the name Linosyris mandoni in Genera Plantarum was its effective place of publication which is clearly incorrect.