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Calea L., Sp. Pl. ed. 2, 2: 1179 (1763).

Leontophthalmum Willd., Ges. Naturf. Fr. Berlin Mag. 1: 140 (1807). Generic description only with no species. Type: Leontophthalmum peruvianum Kunth = Calea peruviana (Kunth) S. F. Blake

Mocinna Lag., Gen. Sp. Pl. Nov. : 31 (1816). Type: Mocinna serrata Lag. = Calea urticifolia (Mill.) DC.

Caleacte R.Br., Trans. Linn. Soc. London 12: 109 (1817), nom. provis. (acc. to Robinson 2006: 75).

Calea L. subgen. Haplocalea Less., Syn. Gen. Comp.: 241 (1832). Type: Calea cymosa Less.

Calea L. subgen. Monanthocalea Less., Syn. Gen. Comp.: 242 (1832). Type: Calea uniflora Less.

Lemmatium DC., Prodr. 5: 670 (1836). Type: Lemmatium rotundifolium (Less.) DC. = Calea rotundifolia (Less.) Baker

Calea L. sect. Amphicalea DC., Prodr. 5: 672 (1836). Type: Calea gentianoides DC.

Meyeria DC., Prodr. 5: 670 (1836). Type not stated. Lectotype (cited by Robinson, 2006: 75): Meyeria myrtifolia DC. = Calea myrtifolia (DC.) Baker

Schomburghia DC., Prodr. 7: 293 (1838). Type: Schomburghia caleoides DC. = Calea caleoides (DC.) H. Rob.

Trinchinettia Endl., Gen. : 1383 (1841), nom. nov. (based on Schomburghia DC.)

Aschenbornia Schauer, Linnaea 19: 16 (1847). Type: Aschenbornia heteropoda Schauer = Calea ternifolia Kunth

Amphicalea (DC.) Gardner, London J. Bot. 7: 411 (1848).

Stenophyllum Sch.Bip. ex Benth. & Hook.f., Gen. Pl. 2: 391 (1873), cited by Robinson (2006: 75) as nom. superfl., although no such name was proposed by Bentham & Hooker f.!

Tonalanthus Brandegee, Univ. Calif. Publ. Bot. 6: 75 (1914). Type: Tonalanthus aurantiacus Brandegee = Calea megacephala B. L. Rob. & Greenm.

Brasilia G. M. Barroso, Arch. Jard. Bot. Rio de Janeiro 17: 19 (1962). Type: Brasilia sickii G. M. Barroso = Calea sickii (G. M. Barroso) Urbatsch, Zlotsky & Pruski.

Calea L. sect. Haplocalea (Less.) Pruski, Kew Bull. 53(3): 683 (1998). Type: Calea cymosa Less.

Calea L. sect. Monanthocalea (Less.) Pruski, Kew Bull. 53(3): 684 (1998).

Note: Panero [2006](2007) also placed the monospecific Venezuelan Tyleropappus Greenm. within the synonymy of Calea.

Type: Santolina jamaicensis L. = Calea jamaicensis (L.) L.


Pruski, J. F. (1984). Calea brittoniana and Calea kristiniae : two new Compositae from Brazil. Brittonia 36(2): 98103.

Pruski, J. F. (1998). Novelties in Calea (Compositae: Heliantheae) from South America. Kew Bull. 53(3): 683693.

Pruski, J. F. & L. E. Urbatsch (1987). Calea dalyi (Compositae: Heliantheae), a new species from the Serrana de Santiago, Bolivia. Brittonia 39(2): 201204.

Urbatsch, L. E., Zlotsky, A. & J. F. Pruski. (1986). Revision of Calea sect. Lemmatium (Asteraceae: Heliantheae) from Brazil. Syst, Bot. 11(4): 501--114.

Wussow, J. R., Urbartsch, L. E. & G. A. Sullivan. (1985). Calea (Asteraceae) in Mexico, Central America, and Jamaica. Syst. Bot. 10(3): 241267.

Calea amellus (L.) L., Sp. Pl., ed. 2 : 1179 (1763) = Salmea scandens (L.) DC.

*Calea anomala Hassl., Feddes Repert. Spec. Nov. Regni Veg. 7: 356 (1909). Type: Bolivia: Im Kamp von Santiago de Chiquitos, ca. 600 m, Mai 1907, leg. Herzog no. 616. Holotype: ?B.

Bolivia (Santa Cruz), Brazil.

Calea aspera Jacq., Collect. 2: 290 (1788) = Melanthera nivea (L.) Small

*Calea brevifolia Rusby, Bull. New York Bot. Gard. 8(No. 28): 132 (1912). Types: [Bolivia:] Apolo, 5000 ft., Apr. 9, 1902 ([R.S. Williams] No. 263)./ The same as Holtons No. 348. Syntypes: NY.

Bolivia (Santa Cruz).

Calea congesta Rusby, Descr. S. Amer. Pl. : 155 (1920) = Calea solidaginea Kunth

*Calea coriacea DC., Prodr. 5: 675 (1836). Type: L in Peruvi legit cl. Haenke ex h. acad. reg. Monac. ... (v.s. in h. supr cit.). Holotype: M; isotype: G-DC (fragm.).

*Calea robusta Britton, Bull. Torrey Bot. Club 19: 151 (1892). Type: Yungas, 4,000 ft. ([Rusby] 2137). Apparently the same as specimens in Herb. Kew collected by R. Pearce at Puente Grande and Quichara; these have somewhat smaller heads. Isosyntype (Rusby 2137): US. Pearce syntypes: K.

Bolivia (Cochabamba, La Paz).

*Calea cymosa Less., Linnaea 5: 158 (1830). Type: not cited other than a reference to (Bras. capit. radiata).

Bolivia (?), Brazil. This is based on a Bang collection s.n., s. loc., listed by Rusby (1907: 390).

Calea dalyi Pruski & Urbatsch, Brittonia 39(2): 201 (1987). Type: BOLIVIA. Santa Cruz. Chiquitos: S slope of the Serrana de Santiago, 510 km E of town of Santiago de Chiquitos, 18˚23'S, 59˚30'W, 800--950 m, 20 Jul 1983, D. C. Daly, M. J. G. Hopkins, L. E. Forero, S. Beck, N. Hernndez, H. Phipps III & H. Wold 2172. Holotype: NY; isotypes: K, LPB, UB, US.

Bolivia (Santa Cruz).

Calea herbert-smithii Rusby, Descr. S. Amer. Pl. : 156 (1920) = Calea solidaginea Kunth

Calea lanceolata Rusby, Bull. New York Bot. Gard. 8(No. 28): 132 (1912) = Oyedaea lanceolata (Rusby) S. F. Blake

Calea lobata (L.) Sw., Prodr. : 113 (1788) = Neurolaena lobata (L.) R.Br. ex Cass.

Calea lutea Pruski & Urbatsch, Brittonia 40(4): 353 (8 Dec. 1988). Type: BRAZIL. Par : Mun. de Itaitubs, savana, arredores da Base Area do Cachimbo, pr ximo ao destacamento km 6 da estrada que vai para o Aeroporto km 794, 922'S, 5455'W, 25 Apr 1983, M. N. Silva, I. L. Amaral, J. Lima, O. P. Monteiro & J. Colho 76. Holotype: INPA; isotypes: F, G, K, LSU, NY, S, UB, US.

Calea saddiana G. M. Barroso, Arch. Jard. Bot. Rio de Janeiro 28: 195 (1986-1987)[Jan. 1989]. Type: ''Brasil, Mato Grosso, Municpio de Santo Ant nio de Leverger, regio da Gruta (Casa de Pedra) de S o Vicente; 13 Jan. 1987, Saddi et al. 7920. Holotype: UFMT; isotypes: RB, UFMT.

Bolivia (Santa Cruz), Brazil.

Calea nematophylla Pruski, Kew Bull. 53(3): 684 (1998). Type: Bolivia. Santa Cruz: Velasco, Parque Noel Kempff M., Campamento La Torre, camino a la parcela, 1339'14"S, 6049'52"W, 300 m, 11 May 1994, Gutirrez, Quevedo & Mamani 899. Holotype: US; isotypes: MO, SCV.

Bolivia (Santa Cruz).

Calea perforata Klatt, Leopoldina 20: 95 (1884) = Calea solidaginea Kunth

*Calea rhombifolia S. F. Blake, Proc. Biol. Soc. Wash. 36: 53 (1923). Type: ''Type in the U.S. National Herbarium, No. 1,120,944, collected on an open pampa at Lake Rogagua, District of Yacuma, Province of Beni, Bolivia, altitude 305 meters, October, 1921, by H. H. Rusby (Mulford Biological Exploration of the Amazon Basin, No. 2164). Holotype: US.

Bolivia (Ben, Santa Cruz).

Wood & Guzman 17472, 27.10.2001 (K)

*Calea robusta Britton, Bull. Torrey Bot. Club 19: 151 (1892) = Calea coriacea DC.

Calea scandens Griseb., Abh. K nigl. Ges. Wiss. Gttingen 24(1): 199 (1879); Symb. Fl. Argent. : 199 (1879) = Calea solidaginea Kunth

*Calea solidaginea Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. 4 (ed. folio): 231 (1818). Type: Crescit in declivitate nobilissimi montis Silla de Caracas, alt. 800 hex. L ? Floret Januario. Holotype: P-Bonpl.

Calea scandens Griseb., Abh. K nigl. Ges. Wiss. Gttingen 24(1): 199 (1879); Symb. Fl. Argent. : 199 (1879). Type: O.: Tarija, Cuesta de Luisina, pr. Salinas. Holotype: Lorentz & Hieronymus 644, GOET.

Calea perforata Klatt, Leopoldina 20: 95 (1884). Type: .'Columbia, in ripa fluminis Mayqueteae, leg. Ed. Otto, Nr. 459. Herbarium Klatt. Holotype: GH.

Calea congesta Rusby, Descr. S. Amer. Pl. : 155 (1920). Types: Specimen sent as part of Herbert H. Smiths No 648 from Colombia, but distinct in every character except the size and form of the heads. No. 2617, from near Masinga Vieja, 1000 feet, December 20, is the same. Of this, Smith says: Trailing or scandent, to 5 or 10 feet, along borders of forest. Previosuly confused with 648, which is a shrub and differs in the flowers. Syntypes: NY.

Calea herbert-smithii Rusby, Descr. S. Amer. Pl. : 156 (1920). Type: A shrub 3 to 5 feet high, common in camps, 1000 to 3000 feet, flowering in December. Specimens from Onaca (Herbert H. Smith, Colombia, No. 648). Holotype: NY.

Bolivia (Chuquisaca, La Paz, Santa Cruz), Brazil, Colombia, French Guiana, Guyana, Suriname, Trinidad, Venezuela.

Calea suriani Cass., Dict. Sci. Nat. 6, suppl. : 33 (1817) = Neurolaena lobata (L.) R.Br. ex Cass.