This page has been extracted from PDF and some formating may have been lost.
To see the original document click here


Ayapana Spach, Hist. Nat. Vg. Phan. 10: 290 (1841).

Type: Ayapana officinalis Spach = Ayapana triplinervis (Vahl) R. M. King & H. Rob.


King, R. M. & H. Robinson. (1970). Studies in the Eupatorieae (Compositae). XXX. The genus Ayapana. Phytologia 20(3): 210212.

King, R. M. & H. Robinson. (1976). Studies in the Eupatorieae (Asteraceae). CLIX. Additions to the genus, Ayapana. Phytologia 34(1): 57--66.

Ayapana amygdalina (Lam.) R. M. King & H. Rob., Phytologia 20(3): 211 (1970).

*Eupatorium amygdalinum Lam., Encycl. 2: 408 (1788). Type: Peru, Cette plante a t trouve au P rou par M. Joseph de Jussieu. (v.s. in h. Juss.). Holotype: P-JU.

Eupatorium loniceroides Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 91 (1818). Type: [Colombia:] Crescit in siccis, temperatis Andium Novo-Granatensium inter Popayan et Pindamon, alt. 950 hex. L Floret Octobri. [Humboldt & Bonpland n. 1914] Holotype: P-Bonpl.

Eupatorium salzmannianum DC., Prodr. 5: 159 (1836). Type: K circa Bahiam Brasiliae in collibus aridis legit cl. Salzmann. [41] ... (v. s.) Holotype: G-DC; isotypes: K 2.

Eupatorium dodonaeifolium DC., Prodr. 5: 161 (1836). Type: L in Peruvi legit cl. Poeppig. ... (v. s. comm. cl. Poeppig sub n. 18.) Holotype: G-DC.

Eupatorium subobtusum DC., Prodr. 5: 161 (1836). Types: L in Brasili circa Bahiam legit cl. Blanchet (n. 1409!) et in siccis ad montem Sineri prope Cayennam cl. Leprieur! ... (v. s.) Syntypes: G-DC.

Eupatorium oxychlaenum DC., Prodr. 5: 162 (1836). Type: ''L in Bras. prov. Minarum Gener. prope Mariannum legit cl. Vauthier (n. 280!). ... (v. s.). Holotype: G-DC.

Eupatorium ixodes Benth., Ann. Nat. Hist. 2(8): 108 (Oct. 1838). Type: Savannahs of the Rupunoony. Schomburgk, n. 79. Holotype: K; isotypes: FI, NY.

Eupatorium ixodes Benth., J. Bot. (Hooker) 2(9): 41 (Feb. 1840), nom. illegit., superfl. Note: this repeats Benthams earlier publication of the name in the Annals of Natural History, q.v.

Eupatorium barclayanum Benth., Bot. Voy. Sulphur: 112 (April 1845). Type: [Panama:] Isle of Taboga, Bay of Panama. [Barclay, s.n.] Holotype: K. [Note: annotations on p. 182 of this work indicate that Mr Hinds types are in Herb. Benth.; Mr Barclays types in Herb. Hook., and; Dr Sinclairs types are in both Herb. Hook. (primarily) and Herb. Benth. (duplicates)]

Bulbostylis (as Bolbostylis) micrantha Gardner, London J. Bot. 6: 449 (1847), nom. nud.

Bulbostylis (as Bolbostylis) elegans Gardner, London J. Bot. 5: 467 (1846). Types: [Brazil:] Hab. Serra da Batalha, district of the Rio Preto, Province of Pernambuco (n. 2899) [NB. This is now considered to be in Bahia State], and in dry Campos near the Rio Claro, Province of Minas Geraes (n. 4843), Brazil. Fl. in June. Syntypes: K (ex herb. Hookerianum). Isosyntype (Gardner 2899): K (ex herb. Benthamianum), NY.

Bulbostylis (as Bolbostylis) tomentosa Gardner, London J. Bot. 5: 468 (1846). Type: [Brazil:] Hab. Elevated Campos in the Diamond District, Brazil. Fl. in July. [Gardner] 4844.

Bulbostylis (as Bolbostylis) microcephala Gardner, London J. Bot. 5: 468 (1846). Type: [Brazil:] Hab. Serra de Araripe, Province of Cear, Brazil. Fl. from Oct. to Dec. [Gardner] 1734. Types: BM, K, US.

Bulbostylis (as Bolbostylis) glandulosa Gardner, London J. Bot. 5: 469 (1846). Type: [Brazil:] ''Hab. Province of Minas Geraes, Brazil. [Gardner] [no number cited]. Gardner (1846: 469) also added I have unfortunately lost the number and exact locality of this plant, but it exists among my Minas Geraes Collections, there can be no doubt that it is from that province.

*Eupatorium loniceroides Kunth var. b minarum Sch.Bip., Linnaea 30: 182 (1859/60), nom. nud.

Eupatorium amygdalinum Lam. var. b elegans (Gardner) Baker in Mart., Fl. Bras. 6(2): 313 (1876).

Eupatorium amygdalinum Lam. var. g dissitiflora Baker in Mart., Fl. Bras. 6(2): 314 (1876). Type: [Brazil:] Ad Rio Bagagem: Pohl n. 281. Holotype: K (ex herb. Benthamianum); isotypes: K, (ex herb. Hookerianum), W. Note: Although Baker saw, and annotated both sheets at K, only the one cited here as the holotype has Rio Bagagem written on the label.

Eupatorium amygdalinum Lam. var. d glandulosa (Gardner) Baker in Mart., Fl. Bras. 6(2): 314 (1876).

Eupatorium glandulosum Sch.Bip. ex Baker in Mart., Fl. Bras. 6(2): 314 (1876), nom. nud. pro syn., non Kunth (1818)(= Ageratina adenophora (Spreng.) R. M. King & H. Rob.)

*Eupatorium amygdalinum Lam. var. e oxychlaenum (DC.) Baker in Mart., Fl. Bras. 6(2): 314 (1876).

Eupatorium glandulosissimum Malme, Kongl. Svenska Vetenskapsakad. Handl. 32(5): 40 (1899). Type: [Brazil:] Matto Grosso: Serra da Chapada, Buriti (In campo aprico, arenoso, graminoso, nebulis saepe humefacto. 1824/694. Malme 1744.). Holotype: S.

Eupatorium hasslerianum Chodat, Bull. Herb. Boiss., ser. 2, 3(8): 711 (1903). Type: Suffrutex 0,3-0,6, petala roseo-violacea in campo Ipe-hu, Sierra de Maracayu, Nov., [Hassler] n. 5279. Holotype: G.

Eupatorium oxychlaenum DC. f. hasslerianum (Chodat) Hassler, Feddes Repert. Spec. Nov. Regni Veg. 11: 175 (1912).

Bolivia (Ben, La Paz, Santa Cruz), Brazil (Bahia, Minas Gerais, S o Paulo), Colombia, Costa Rica, Ecuador, Guatemala, Guayana, Honduras, Nicaragua, Panama, Peru, Paraguay, Trinidad, Venezuela.

Guko (Cabrera, 1996).

Ayapana lanceolata R. M. King & H. Rob. Phytologia, 34(1): 59 (1976).Type: PERU: San Martin: Prov. & Dist. Lamas, north of San Antonio 2'-4 km, along Ro Cumbasa in dense jungle. Alt. ca. 1200 ft. Vine to 10 feet; flowers probably white. Oct. 2- Nov. 4, 1937. Belshaw 3517. Holotype: US. Amongst the paratypes King & Robinson (1976: 60) cited: ''BOLIVIA: La Paz: Region tropical, San Carlos b. Mapiri, 750 m, in Gebsch, Strauch 1 m hoch. August 1907. Buchtien 1507 (US); Prov. of S. Yungas, basin of Rio Bopi, San Bartolome (near Calisaya), alt. 75900 m. Herb. July 122, 1939. Krukoff 10353 (US).

Bolivia(La Paz), Peru.

Ayapana pyramidalis (Klatt) R. M. King & H. Rob., Phytologia 20(3): 212 (1970) = Ayapana stenolepis (Steetz) R. M. King & H. Rob.

Ayapana stenolepis (Steetz) R. M. King & H. Rob., Phytologia 32(3): 284 (1975).

Eupatorium stenolepis Steetz in Seem., Bot. Voy. Herald : 148 (1854). Type: Santiago de Veraguas. [March 1848, Seemann 1135]'' Holotype: BM; isotype: K.

Eupatorium amygdalinum var. z revoluta Baker in Mart., Fl. Bras. 6(2): 314 (1876). Type: [Brazil:] Inter Rio Abait et Apollonario: Pohl n. 500. Holotype: K (ex herb. Benthamianum); isotypes: K (ex herb. Hookerianum), W.

Eupatorium revolutum Pohl ex Baker in Mart., Fl. Bras. 6(2): 314 (1876), nom. nud. pro syn.

Eupatorium rhodanthum Sch.Bip. ex Baker in Mart., Fl. Bras. 6(2): 314 (1876), nom. nud. pro syn.

*Eupatorium pyramidale Klatt, Abh. Naturf. Ges. Halle 15: 323 (1881), nom. illegit., non D. Don (1825) (= Vernonia aspera (Roxb.) Ham.[VERNONIEAE]. [Type (from separate in K, on p. 1): ''Crescit in Provincia Chiquito, leg. A. dOrbigny No. 680.] According to Robinson (1920) isotype: GH. Note: Although Klatts herbarium was purchased by GH, Herb. Francaville is in P; isotype: G. [See note under Antennaria linearifolia]

*Eupatorium amygdalinum Lam. var. z revolutum Baker f. angustifolium Hieron. ex Kuntze, Revis. Gen. Pl. 3(3): 146 (1898), nom. nud.

*Eupatorium pyramidale Klatt f. b angustifolium B. L. Rob., Contr. Gray Herb. 61: 67 (1920). Type: [Bolivia:] Santa Cruz: Prov. East Velasco, alt. 200 m., Kuntze. Holotype: NY.

Ayapana pyramidalis (Klatt) R. M. King & H. Rob., Phyotlogia 20(3): 212 (1970).

Bolivia (Santa Cruz), Panama.