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Werneria Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 148 (1818).

Euryops Cass., Bull. Sci. Soc. Philom. Paris 1818: 141 (1818).

Oresigonia Willd. ex DC., Prodr. 6: 323 (1838), nom. nud. pro syn.

Oribasia Mo. & Sesse ex DC., Prodr. 6: 323 (1838), nom. nud. pro syn.

Wernera Kuntze, Revis. Gen. Pl. 3(3): 184 (1898), orth. var. (but only as part of a binomial)

Werneria Kunth subgen. Anactis (Gay) Rockh., Bot. Jahrb. Syst. 70(3): (1939), comb. illegit.

Werneria Kunth subgen. Anactis (Gay) Rockh. sect. Pinnatifoliae Rockh., Bot. Jahrb. Syst. 70(3): (1939), nom. illegit.

Werneria Kunth subgen. Anactis (Gay) Rockh. sect. Integrifoliae Rockh., Bot. Jahrb. Syst. 70(3): (1939), nom. illegit.

Werneria Kunth subgen. Euwerneria (Gay) Rockh., Bot. Jahrb. Syst. 70(3): (1939), comb. illegit.

Werneria Kunth subgen. Euwerneria (Gay) Rockh. sect. Graminifoliae Rockh., Bot. Jahrb. Syst. 70(3): (1939), nom. illegit.

Werneria Kunth subgen. Euwerneria (Gay) Rockh. sect. Spathulifoliae Rockh., Bot. Jahrb. Syst. 70(3): (1939), nom. illegit.

Note: Rockhausens account (1939) provided a number of illegitimate infrageneric names (as only the name was proposed, lacking any Latin description, contrary to Art. 36 of the present ICBN, 2006).

Type: not stated. Lectotype (?selected by Cabrera, 1971: 275): Werneria graminifolia Kunth. Note: Funk (1997b) indicated that the type was W. nubigena Kunth although I have been unable to find any reference that lectotypified the genus based on this name.

References

Cabrera, A. L. (1948). Las especies Argentinas del g nero Werneria. Notas Mus. La Plata, Bot. 13(No. 60): 4961.

Blake, S. F. (1928). New South American species of Werneria. J. Wash. Acad. Sci. 18(18): 485498.

Funk, V. A. (1997). Werneria s.l. (Compositae: Senecioneae) in Ecuador. In: Valencia, R. & H. Balslev (eds), Estudios sobre diversidad y ecologia de plantas. Memorias del II Congreso Ecuatoriano de Botnica. Univ. Cat l. Quito: 2535.

Rockhausen, M. (1940). Vervandtschaft und Gliederung der Compositen-Gattung Werneria. Bot. Jahrb. Syst. 70(3): 273342.

Werneria apiculata Sch.Bip., Bonplandia 4(4): 52 (1856) = Werneria pygmaea Gillies ex Hook. & Arn.

*Werneria aretioides Wedd., Chloris Andina 1: 86 (1856). Type: Hab. Bolivie: sur les montagnes des lagunas de Potosi! (dOrbigny, no 1400). Holotype: P.

Argentina, Bolivia (Potos), Chile, Peru.

Werneria articulata S. F. Blake, Contr. U. S. Natl. Herb. 22: 651 (1924), nom. nov. pro W. lehmannii Hieron. = Xenophyllum humile (Kunth) V. A. Funk

*Werneria boraginifolia Kuntze, Revis. Gen. Pl. 3(3): 184 (1898) = Misbrookea strigosissima (A. Gray) V. A. Funk

Werneria brachypappus Sch.Bip., Bonplandia 4(4): 53 (1856), non Phil. (1873) (= Werneria denticulata S. F. Blake following Rockhausen, although not treated by him) = Werneria pygmaea Gillies ex Hook. & Arn.

*Werneria caespitosa var. b haenkei Wedd., Chloris Andina 1: 83 (1856). Although listed by Foster (1958), Rockhausen (1939: 316) suggested that this is only a Peruvian taxon, although it is likely that the Bolivian record was based on Troll 2130; Rockhausen only recognized the species.

*Werneria caulescens (Wedd.) Rusby, Bull. New York Bot. Gard. 4(14): 398 (1907) = Werneria villosa A. Gray

*Werneria ciliata Wedd. ex Sch.Bip., Linnaea 34(5): 530 (Feb. 1866), pro syn. = Werneria pectinata Lingelsh.

*Werneria ciliolata A. Gray, Proc. Amer. Acad. Arts 5: 140 (1861) = Xenophyllum ciliolatum (A. Gray) V. A. Funk

Werneria cortusifolia Griseb., Abh. K nigl. Ges. Wiss. Gttingen 19([1]): 194 (1874); Pl. Lorent. : 146 (1874) = Senecio breviscapus DC.

*Werneria dactylophylla Sch.Bip., Bonplandia 4(4): 52 (1856) = Xenophyllum dactyophyllum (Sch.Bip.) V. A. Funk

*Werneria dactylophylla Sch.Bip. f. glabriuscula Rockh., Bot. Jahrb. Syst. 70: 286 (1939) = Xenophyllum dactyophyllum (Sch.Bip.) V. A. Funk

Werneria dactylophylla Sch.Bip. var. glanduloso-denticulata Rockh., Bot. Jahrb. Syst. 70(3): 286 (1939) = Xenophyllum dactyophyllum (Sch.Bip.) V. A. Funk

?Werneria decumbens Hieron., Bot. Jahrb. Syst. 21(4): 364 (1896) = Xenophyllum weddellii (Phil.) V. A. Funk

*Werneria digitata Wedd., Chloris Andina 1: 86 (1856) = Xenophyllum digitatum (Wedd.) V. A. Funk

Werneria digitata Wedd. var. lanata Rockh., Bot. Jahrb. Syst. 70(3): 287 (1939) = Xenophyllum digitatum (Wedd.) V. A. Funk

*Werneria heteroloba Wedd., Chloris Andina 1: 88 (1856). Types: Hab. P rou: Cordillres du d partement de Cuzco! (Gay). Bolivie: sur les pelouses marcageuses de la Lancha!, Cordill re de La Paz, avec le W. pygmaea (Wedd.); bord des eaux, dans les montagnes des lagunas de Potosi! (dOrbigny, no 1415). Syntypes: P.

Werneria obtusiloba S. F. Blake, J. Wash. Acad. Sci. 18(18): 489 (1928). Type: Peru: In sandy soil, with cushion and rosette plants, cordillera east of Carumas, Prov. Moquegua, alt. 4500-4600 m., 7-8 Mar. 1925, A. Weberbauer 7362. Holotype: F (552591); isotype: US (44298).

Werneria heteroloba Wedd. var. obtusiloba (S. F. Blake) Rockh., Bot. Jahrb. Syst. 70(3): 283 (1939).

Werneria heteroloba Wedd. f. microcephala Rockh., Bot. Jahrb. Syst. 70: 283 (1939). Type: Chile: Lago Chungara, in Smpfen, 4550 m .M. (C. Troll Nr. 3222 Mrz 1927.).

Argentina, Bolivia (Potos ), Chile, Peru.

Werneria heteroloba Wedd. f. microcephala Rockh., Bot. Jahrb. Syst. 70: 283 (1939) = Werneria heteroloba Wedd.

Werneria heteroloba Wedd. var. obtusiloba (S. F. Blake) Rockh., Bot. Jahrb. Syst. 70(3): 283 (1939) = Werneria heteroloba Wedd.

Werneria humilis Kunth in Humb., Bonpl. & Kunth., Nov. Gen. Sp. Pl. 4 (ed. folio): 150 (1818) = Xenophyllum humile (Kunth) V. A. Funk

Werneria humilis Kunth f. articulata (S. F. Blake) Rockh., Bot. Jahrb. Syst. 70(3): 294 (1939) = Xenophyllum humile (Kunth) V. A. Funk

?Werneria juniperina Hieron., Bot. Jahrb. Syst. 21(4): 365 (1896) = Xenophyllum ciliolatum (A. Gray) V. A. Funk

*Wereneria knocheae Perkins, Bot. Jahrb. Syst. 49: 230 (1913) = Werneria pectinata Lingelsh.

Werneria lehmannii Hieron., Bot. Jahrb. Syst. 28(5): 647 (1901), non Klatt (1894) = Xenophyllum humile (Kunth) V. A. Funk

Werneria lorentziana Hieron., Bot. Jahrb. Syst. 21: 364 (1896) = Xenophyllum poposum (Phil.) V. A. Funk

Werneria macbridei Cuatrec., Collect. Bot. (Barcelona) 3(3): 294 (1953) = Senecio repens DC. var. macbridei (Cuatrec.) Cabrera

*Werneria mandoniana Wedd. ex Klatt, Ann. K. K. Naturh. Hofmus. Wien 9: 367 (1894) = Werneria orbignyana Wedd.

Werneria marcida S. F. Blake, J. Wash. Acad. Sci. 18(18): 492 (1928) = Xenophyllum marcidum (S. F. Blake) V. A. Funk

*Werneria melanandra Wedd., Chloris Andina 1: 88 (1856). Types: Hab. Bolivie: province de Carangas (dOrbigny, no 1395); pelouses humides de la Lancha!, dans la partie suprieure du ravin de Chuquiguillo, pr s de La Paz (Wedd.). Syntypes: P.

Bolivia (La Paz), Peru.

Werneria mociniana DC., Prodr. 6: 324 (1838) = Werneria nubigena Kunth

*Werneria nubigena Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 151 (1818). Type: [Ecuador:] Crescit in frigidis montis Chimborazi, alt. 1700 hex. (Regno Quitensi.) K Floret Junio. [Humboldt & Bonpland P-Bonpl - 3191. Chimborazo; B-W: 3191] Holotype: P-Bonpl; isotype: B-W.

Werneria disticha Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): (1818). Type: Crescit in summo monte ignivomo Antisan, alt. 2100 hex. K Floret Junio. [Humboldt & Bonpland 2272] Holotype: P-Bonpl; isotype: B-W.

B: ć¤‰í˛‘2272. Antisana

Oresigonia latifolia Willd. ex DC., Prodr. 6: 324 (1838), nom. nud. pro syn.

Werneria mociniana DC., Prodr. 6: 324 (1838). Type: K in Mexico ad cacumen montis cuchumatanes. Orisaba acaulis Moc. Sess. et Cerv. incon fl. mex. ined. ... (v. ic. pict.)

Oribasia acaulis Moc., Sesse & Cerv. ex DC., Prodr. 6: 324 (1838), nom. nud. pro syn.

Werneria nubigena Kunth var. a vulgaris Wedd., Chloris Andina 1: 80 (1855).

Werneria nubignea Kunth var. b latifolia Wedd., Chloris Andina 1: 80 (1855), nom. illegit. pro Werneria disticha Kunth

Werneria steubelii [as stbelii] Hieron., Bot. Jahrb. Syst. 21(4): 362 (1896). Type: Peruvia: crescit prope Challuayacu inter Pacasmayo et Moyobamba, alt. s. m. 3400 m, ubi floret mense Aprili-Junio ([St bel] coll. peruv. n. 55c).

Oresigonia grandiflora Willd. ex Rockh., Bot. Jahrb. Syst. 70(3): 301 (1939), nom. nud. pro syn.

Bolivia (Cochabamba, La Paz), Colombia, Ecuador, Guatemala, southern Mexico, Peru.

Werneria nubignea Kunth var. b latifolia Wedd., Chloris Andina 1: 80 (1855), nom. illegit. pro Werneria disticha Kunth= Werneria nubigena Kunth

Werneria nubigena Kunth var. a vulgaris Wedd., Chloris Andina 1: 80 (1855) = Werneria nubigena Kunth

Werneria obtusiloba S. F. Blake, J. Wash. Acad. Sci. 18(18): 489 (1928) = Werneria heteroloba Wedd.

*Werneria orbignyana Wedd., Chloris Andina 1: 85 (1856). Type: Hab. Bolivie: au sommet de la Cordillre de La Paz! (dOrbigny, no 338).č’’

*Werneria mandoniana Wedd. ex Klatt, Ann. K. K. Naturh. Hofmus. Wien 9: 367 (1894). Type: [Bolivia:] Hab.: Prov. Larecaja, viciniis Sorata, valle inter las trincheras de Chiliata et montem Hlampie, in scopulosis, Alt. Reg. alpina, 3800 m., leg. Mandon, Sept. 1858, Nr. 102. Holotype: W; isotype: S.

Bolivia (La Paz), Peru.

*Werneria pectinata Lingelsh., Feddes Repert. Spec. Nov. Regni Veg. 8: 6 (1910). Type: Bolivien: Am Chacaltaya, 4800 m (Buchtien, no. 1596!). Holotype: ; isotype: US.

*Werneria ciliata Wedd. ex Sch.Bip., Linnaea 34(5): 530 (Feb. 1866), nom. nud. pro syn. (based on Mandon 99)

*Wereneria knocheae Perkins, Bot. Jahrb. Syst. 49: 230 (1913). Types: Bolivien: Aguila, 17 sdl. Breite, 67 westl. L nge, an der Cordillera Real, 5200 m . M. (Edith Knoche n. 2, 14, 21. Im April 1909 bl hend). Syntypes: B

Bolivia (La Paz), Peru.

*Werneria plantaginifolia Wedd. ex Klatt, Ann. K. K. Naturh. Hofmus. Wien 9: 367 (1894). Type: [Bolivia:] Hab.: Prov. Larecaja, vinciniis Sorata, prope las trincheras de Chiliata, in graminosis, Reg. alpina, 3800 m., leg. Mandon, Mart.-Mai 1859, Nr. 89. Holotype: W; isotype: S.

Bolivia (La Paz), Peru.

Werneria poposa Phil., Anales Mus. Nac. Chile, Bot. 8: 40 (1891) = Xenophyllum poposum (Phil.) V. A. Funk

*Werneria pygmaea Gillies ex Hook. & Arn., J. Bot. (Hooker) 3: 348 (1841). Type: Valle de los Ciegos, Andes of Mendoza; Dr. Gillies.

Werneria rhizoma Remy in Gay, Hist. Chile, Bot. 4: 215 (1849). Type: [Chile: ] Se halla en las cordilleras de Ovalle, los Patos. Holotype: P.

*Werneria apiculata Sch.Bip., Bonplandia 4(4): 52 (1856). Type: á’‘Meine Werneria apiculata kommy unter 2 Nummern von sterilen Andenweiden vor. [Note: In a later list of determinations it is clear that these were Lechler 1737 and 1964 from Peru. Inference from Schultz Bipontinus paper (Schultz Bipontinus, 1856) is that these were collected between May (cf. Lechler 1710 Werneria brachypappa) and June 1854 (cf. Lechler 1963 Achyropappus cryptocephalus).] [isosyntype (Lechler 1737): S]

Werneria pygmaea Gillies ex Hook. & Arn. var. e apiculata (Sch.Bip.) Wedd., Chloris Andina 1: 84 (1856).

Werneria brachypappus Sch.Bip., Bonplandia 4(5): 53 (1856), non Phil. (1873). Type: [Peru:] [Lechler] 1710a, in uliginosis Cord. de Tuno, Mai 1854 in Cherleria hnlichen Rasen, ... Holotype: ?P.

Argentina, Bolivia (Cochabamba, La Paz, Potos ), Chile, Colombia, Ecuador, Peru,Venezuela.

Werneria pygmaea Gillies ex Hook. & Arn. e apiculata (Sch.Bip.) Wedd., Chloris Andina 1: 84 (1856) = Werneria pygmaea Gillies ex Hook. & Arn.

Werneria rhizoma Remy in Gay, Hist. Chile, Bot. 4: 215 (1849) = Werneria pygmaea Gillies ex Hook. & Arn.

Werneria rosenii R. E. Fr., Nova Acta Regiae Soc. Sci. Upsal., ser. 4, 1(1): 90 (1905) = Xenophyllum rosenii (R. E. Fr.) V. A. Funk

Werneria sedoides S. F. Blake, J. Wash. Acad. Sci. 18(18): 493 (1928) = Xenophyllum marcidum (S. F. Blake) V. A. Funk

*Werneria setosa Wedd. ex Sch.Bip., Linnaea 34(5): 530 (Feb. 1866), nom. nud. pro syn. = Misbrookea strigosissima (A. Gray) V. A. Funk

*Werneria solivifolia Sch.Bip., Bonplandia 4(4): 53 (1856). Type: [Peru:] Nun kimmen noch zwei neue Wernerien zur Aburtheilungm welche mir unter [Lehmann] Nr. 1710, in uliginosis Cord. de Tuno, Mai 1854 in Cherleria hnlichen Rasen, in welchen die kleinen K pfchen versenkt waren, wachsend, mitgetheilt wurden. Note: In a subsequent listing it appears that Werneria brachypappa was based on 1710a, and W. solivaefolia.

Bolivia (La Paz, Potos), Peru.

*Werneria spathulata Wedd., Chloris Andina 1: 85 (1856). Type: Hab. Bolivie: partie sup rieure du ravin de Chuquiaguillo, dans les pelouses un peu tourbeuses de la Lancha!, Cordillre de La Paz (Wedd.). Holotype: P.

Argentina, Bolivia (La Paz), Chile, Peru.

Werneria staffordiae Sandwith, Hookerč’’s Ic. Pl., ser. 5, 5 (vol. 35 of whole work): tab. 3424 pp. 12, (1950) = Xenophyllum staffordiae (Sandwith) V. A. Funk

Werneria steubelii [as stbelii] Hieron., Bot. Jahrb. Syst. 21(4): 362 (1896) = Werneria nubigena Kunth

*Werneria staticifolia Sch.Bip., Bonplandia 4(4): 53 (1856). Type: [Peru: ] St. Gavan in summis Cordil. jugis Jul. 1854: Lechler! Nr. 2212. Holotype: ?P.

Bolivia (La Paz), Peru.

*Werneria strigosissima A. Gray, Proc. Amer. Acad. Arts 5: 140 (1861) = Misbrookea strigosissima (A. Gray) V. A. Funk

*Werneria villosa A. Gray, Proc. Amer. Acad. Arts 5: 139 (1861). Type: High Andes of Peru near Alpamarca [Collections of the United States South Pacific Exploring Expedition under Captain Wilkes]. Holotype: GH.

*Werneria caulescens Griseb., Abh. K nigl. Ges. Wiss. Gttingen 24(1): 208 (March-April 1879); Symb. Fl. Argent. : 208 (1879). Types: Syn. W. nubigena var. caulescens Wedd. ... S.: Nevado del Castillo. (And. boliv.) Note: Weddell (1856: 81) only indicated that four of his varieties of Wereneria nubigena were represented by collections of Gay and Dombey from Peru. Bolivian collections listed, Weddell (3952, 4750) and dí¸…í˛’Orbigny (336), were not related to specific varieties. Two subvarieties, leioscapa and erioscapa were also listed under var. d caulescens no type material was indicated.

Argentina, Bolivia (La Paz), Peru.

Werneria weddellii Phil., Anales Mus. Nac. Chile, Bot. 8: 40 (1891) = Xenophyllum weddellii (Phil.) V. A. Funk

Werneria wernerioides (Wedd.) Kuntze, Revis. Gen. Pl. 3(3): 184 (1898) = Senecio breviscapus DC.