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Verbesina L., Sp. Pl. : 901 (1753)., Gen. Pl. ed. 5: 384 (1754).

Ridan Adans., Fam. Pl. 2: 120, 598x (1763), nom. rej. Actinomeris Nutt. (1818), nom. cons. Type: Coreopsis alternifolia L. Verbesina alternifolia (L.) Britt. ex Kearney

Phaethusa Gaertn., Fruct. Sem. Pl. 2: 425, pl. 169 (1791). Type: Phaethusa americana Gaertn. = Verbesina occidentalis (L.) Walt.

Ximenesia Cav., Icon. 2: 60, pl. 178 (17931794). Type: Ximenesia encelioides Cav. = Verbesina encelioides (Cav.) Benth. & Hook.f. ex A. Gray

Hingstonia Raf., Med. Repos. N. York 5: 352 (1808). Hingstonia exalata Raf. = Verbesina occidentalis (L.) Walt.

Ditrichum Cass., Bull. Sci. Soc. Philom. Paris 1817: 33 (1817), nom. rej., non Hampe. Type: Ditrichum macrophyllum Cass. = Verbesina macrophylla (Cass.) S. F. Blake

Actinomeris Nutt., Gen. 2: 181 (1818), nom. cons. vs. Ridan Adans. Type: Actinomeris squarrosa Nutt., nom. illeg. (Coreopsis alternifolia L.) type cons. = Verbesina alternifolia (L.) Britt. ex Kearney

Platypteris Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 156 (1818). Type: Bidens crocata Cav. = Verbesina crocata (Cav.) Less.

Actimeris Raf., Amer. Monthly Mag. & Crit. Rev. : 195 (1819) Actinomeris nutt.

Hamulium Cass., Bull. Sci. Soc. Philom. Paris 1820: 173 (1820). Type: Verbesina alata L.

Ochronelis Raf., Atlantic J.: 153 (1832). Type: Ochronelis sulfurea Raf. = Verbesina aristata (Ell.) A.Heller

Saubinetia Remy in Gay, Hist. Chile Bot. 4: 282, pl. 49 (1849). Type: Saubinetia helianthoides Remy = Verbesina saubinetia Klatt

Ancistrophora A. Gray, Mem. Amer. Acad. 2, 6: 457 (1859). Type: Ancistrophora wrightii A. Gray = Verbesina lindheimeri B. L. Rob. & Greenm.

Chaenocephalus Griseb., Fl. Brit. W. I. : 374 (1861). Type: Chaenocephalus petrobioides Griseb. Verbesina petrobioides (Griseb.) S. F. Blake

Achaenopodium Brand., Zoe 5: 239 (1906). Type: Achaenopodium discoideum Brand. = Verbesina discoidea (Brand.) Rzed.

Lectotype: Verbesina alata L.

References

Blake, S. F. (1925). On the status of the genus Chaenocephalus, with a review of the section Lipactinia of Verbesina. Amer. J. Bot. 12(10): 625640.

Coleman, J. R. (1966). A taxonomic revision of Section Ximenesia of the genus Verbesina L. (Compositae). Amer. Midl. Nat. 76(2): 475481.

Espinar, L. A. (1991). Rehabilitacion de Verbesina aurita (Asteraceae). Kurtziana 21: 279282.

Jeffrey C. (1992). (1058) Proposal to conserve Verbesina L. (Compositae) with a conserved type. Taxon 41(3): 595.

Robinson, B. L. & J. M. Greenman. (1899). Synopsis of the genus Verbesina, with an analytical key to the species. Proc. Amer. Acad Arts 34(20): 534566.

Verbesina alba L., Sp. Pl.: 902 (1753) = Eclipta prostrata (L.) L.

Verbesina allophylla S. F. Blake, Amer. J. Bot. 12(10): 633 (1925), as nom. nov. pro Chaenocephalus heterophyllus Griseb. non Verbesina heterophylla (Chapm.) A. Gray = Verbesina suncho (Griseb.) S. F. Blake

*Verbesina australis Baker in Mart., Fl. Bras. 6(3): 215 (1884), nom. nov. pro Ximenesia microptera DC. = Verbesina encelioides (Cav.) A. Gray

*Verbesina benderi Perkins, Bot. Jahrb. Syst. 49: 227 (1913). Type: Bolivien: Quechisla, 2030' sdl. Breite, 6620' westl. L nge, Schieferboden, 3425 m . M. (C. Bender n. 7. Im M rz 1909 blhend). Holotype: B.

Bolivia (?).

*Verbesina boliviana Klatt, Ann. K. K. Naturhist. Hofmus. Wien 9: 361 (1894). Type: Hab.: Bolivia, leg. Cuming. Holotype: W; GH (13497 - possibly type fragment and drawing).

Verbesina bridgesii Rusby, Mem. Torrey Bot. Club 4(3): 212 (1895). Types: [Bolivia:] á''[Bang] 974. Collected also by Bridges in Bolivia. Holotype: NY; isotype: US.

Bolivia (?).

[Section 9 Saubinetia (Remy) B. L. Rob. & Greenm.] Apparently leaves sometimes lobed in this species.

Verbesina brasiliana Pers., Syn. Pl. 2: 472 (1807) = Adenostemma brasilianum (Pers.) Cass.

*Verbesina bridgesii Rusby, Mem. Torrey Bot. Club 4(3): 212 (1895) = Verbesina boliviana Klatt

Verbesina carnosa (Rich.) G.Maza, Periant. : 274 (1890), nom. illeg. = Sphagneticola trilobata (L.) Pruski

*Verbesina cinerea Rusby, Mem. Torrey Bot. Club 6(1): 63 (1896). Type: [Bolivia:] Below Cochabamba, 1891 ([Bang] 1092). Holotype: NY; isotype: US.

Bolivia (Cochabamba).

[Section 9 Saubinetia DC.]

Verbesina conyzoides Trew, Pl. Rar. : 8 (1763) = Eclipta prostrata (L.) L.

Verbesina cumingii Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea 34(5): 528 (Feb. 1866), nom. nud. (based on Mandon 56 and Cuming 108 or 1108 q.v. Bull. Soc. Bot. France 12: 79 (1865)) = Verbesina cumingii (Griseb.) S. F. Blake

*Verbesina cumingii (Griseb.) S. F. Blake, Amer. J. Bot. 12: 634 (1925).

Verbesina cumingii Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea 34(5): 528 (Feb. 1866), nom. nud. (based on Mandon 56 and Cuming 108 or 1108 q.v. Bull. Soc. Bot. France 12: 79).

Chaenocephalus cumingii Griseb., Abh. Knigl. Ges. Wiss. G ttingen 24(1): 196 (March-April 1879); Symb. Fl. Argent. : 196 (1879). Type: (Verbesinae Sch. in Mand.[on] pl. boliv. 56.) Holotype: GOET; isotypes: GH, NY, S, US (fragment of GH material).[Note: this is a hyponym according to Blake (1925: 634), but perfectly validly described, since it was compared with Chaenocephalus macrophyllus described at that point. The type cited by Blake is the same Mandon collection, but with a fuller locality, albeit in the wrong country: Ecuador [sic! = Bolivia]: in woods, near Sorat, alt. 2800-3100 m., July-August 1858-1859, Mandon 56 (Type in Gray Herb.; duplicate in herb. N. Y. Bot. Gard.; photogr. and fragm., U.S. Nat. Herb.).]

Bolivia (La Paz).

Verbesina diversifolia DC., Prodr. 5: 615 (1836) = Verbesina macrophylla (Cass.) S. F. Blake

*Verbesina elegans Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 160 (1818). Type: [Ecuador:] íYTí²'Crescit in Regno Quitensi ? K ? Holotype: P-Bonpl.

Bolivia (?), Peru.

[Section 8. Verbesinaria DC.]

Verbesina encelioides (Cav.) A. Gray, Bot. California 1: 350 (1876).

Ximenesia encelioides Cav., Icon. 2: 60, t. 178 (1793). Type: Habitat in Mexico unde introducta in Regium hortum Matritensem. Floruit prima vice mense Novembri: videtur perennis. Note: There are four sheets in MA, MA (476510 Fiche 80/C8) with a handwritten label Ximenesia encelioides Cav./ex Bonariensi planitie/Ne Iter., MA (476508 é'- Fiche 81/A1) has three handwritten labels, the upper Ximenesia encelioides/Cav. Ic./ta. 178/Willd. Syng. pa. 2886/ex Hort. Reg. Matr./anno 1808, the lower left Ximenesia encelioides/Cav. Ic. 2 ta. 178/X. pachyptera DC. Prod./Pallasia serratifolia Smith/Ex H. M. Aug. 1844 and the middle label Genus Encelia affine/Ximenesia encelioides/Icon. tab. 178/Mexico Culta in R. h. M.1792-93., MA (476508 sheet 2 Fiche 81/A2) merely has a printed label, and MA (476509 Fiche 81/A3) has two handwritten labels, the upper Ximenesia encelioides/Cav. Ic./Ex H. M. Aug. 1848 the lower Ximenesia enceli/odes [sic!] Cav. Ic. ta. 178/Willd. Syng. pa. 2886/ex Hort. Reg. Matr./anno 1808. This suggests material was continually grown in the botanic garden and specimens prepared and incorporated in the herbarium. Two of these sheets might be considered as type material

Ximenesia encelioides Cav. var. a hortensis DC., Prodr. 5: 627 (1836). Type: Frequens in hortis botanicis sed nunqum specimen cert spontaneum vidi. (v.v.c.)

Ximenesia encelioides Cav. var. b pachyptera DC., Prodr. 5: 627 (1836). Types: in Mexico legit cl. Berlandier (pl. exs. n. 2068 et 2286!). (v.s.sp.). Note: I do not consider Colemans citation of a single Berlandier specimen P? as effective lectotypification (cf. Coleman, 1966: 476).

Ximenesia encelioides Cav. var. g oblongifolia DC., Prodr. 5: 627 (1836). Types: in Senegali ad ripas flum. regionis Walo legit Perrottet, in insulis Mauritianis Sieber (fl. maur. 2. n. 118!) et Belanger, et in hort. Calcuttensi Wallich (v.s.)

Ximenesia encelioides Cav. var. d? cana DC., Prodr. 5: 627 (1836). Types: â''in Mexico circa Laredo legit cl. Berlandier (pl. exs. n. 2068 et 2074). ...v.s.). Note: I do not consider Colemans selection of a single Berlandier specimen P? as effective lectotypification.

Ximenesia microptera DC., Prodr. 5: 627 (1836). Type: circa Buenos-Ayres legit cl. Bacle [32]. ... (v.s.) Holotype: G-DC.

Ximenesia australis Hook. & Arn. ex DC., Prodr. 7: 291 (1838), nom. nud. pro syn. Note: Coleman (1966: 477) both cited erroneous authorities (Benth. & Arn. in DC.) and assumed there was a type for this name.

Verbesina scabra Phil., Anales Univ. Chile 36: 186 (1870), non Benth. (1840). [Note: In a separately paginated reprint/preprint of Sertum Mendocinum alterum, pp. 154, this appears on p. 28.] Type: Mendoza. Pizarro (1960: 165) cited SGO 44007 and 65406.

Encelia (Geraea) albescens A. Gray, Proc. Amer. Acad. Arts 8: 658 (1873). Type: In the western Mexican province of Sonora, Dr. Edward Palmer, coll. 1869, no. 21. Lectotype (selected by Coleman, 1966: 477): Sonora, Mexico. Palmer 21 GH; isolectotype: US.

*Verbesina australis Baker in Mart., Fl. Bras. 6(3): 215 (1884), nom. nov. pro Ximenesia microptera DC.

Verbesina encelioides (Cav.) A. Gray var. cana (DC.) B. L. Rob. & Greenm., Proc. Amer. Acad. Arts 34: 544 (1899), p.p.

Verbesina microptera (DC.) Herter, Rev. Sudam. Bot. 7: 235 (1843), comb. illegit. non Verbesina microptera DC. (1836).

Native distribution apparently the Gulf Coast plains of Texas and Mexico. Widely distributed as a pantropic weed as well as frequently cultivated, especially in temperate regions. Bolivia (Chuquisaca, Cochabamba, Santa Cruz).

Note: Several references (Gray, 1876; Coleman, 1966; Hind, 1993; Index Kewensis; etc.) quote the authority of this combination as variously (Cav.) Benth. & Hook.f. ex A. Gray or (Cav.) Benth. ex A. Gray; neither of these are correct. Bentham & Hooker f. provided nothing other than an inference that there were two species of Verbesina sect. Ximenesia no names were mentioned. Since Gray (1876: 350) was actually the first author to validly make the combination it is to him alone it should be attributed. Espinar (1991: 280) has provided useful evidence that Verbesina aurita Phil. (Anales Mus. Nac. Chile, Secc. 2, Bot. 8: 48 (1891)) should be removed from the synonymy of V. encelioides; V. aurita is now considered as restricted to northern Chile and the provinces of Catamarca, Salta and Tucumn in Argentina.

Some authors considered that Baker (1884: 215) was describing a new species of Verbesina. Clearly he was not since he cited Ximenesia microptera DC. in synonymy but, since de Candolle had already described Verbesina microptera Bakers name can be considered as a nom. nov. when X. microptera is recognized as a separate species in Verbesina.

*Verbesina flavovirens R. E. Fr., Arkiv. Bot. 5(13): 19 (1906). Type: á''Bolivia: Pinos pr. Tarija, in fruticetis regionis Alni ferrugineae, ca. 2500 m. s. m. [¹/₃ 02; Fr. 1292]. Holotype: S.

Bolivia (Tarija).

Verbesina foliacea Spreng., Syst. Veg., ed. 16, 3: 578 (1826) = Eleuthernathera ruderalis (Sw.) Sch.Bip.

Verbesina lancifolia Gardner, London J. Bot. 7: 406 (1848) = Verbesina macrophylla (Cass.) S. F. Blake

Verbesina lapazii J. L. Panero, Contrib. Univ. Mich. Herb., 19: 183 (1993). Type: BOLIVIA. La Paz: Prov. Nor Yungas, de Coroico 12 km hacia Coripata, 2050 m, 11 Apr 1990, Beck 17629. Holotype: US; isotype: LPB.

Bolivia (La Paz).

*Verbesina leucactinota B. L. Rob., Proc. Amer. Acad. Arts 47: 213 (1911). Type: Coripati, Yungas, Bolivia, April, 1894, Bang, no. 2135. Holotype: GH; isotypes: US.

Bolivia (La Paz).

Verbesina linifolia L., Syst. Nat., ed. 10, 2: 1226 (1759), nom. superfl. = Pectis linifolia L.

*Verbesina macrophylla (Cass.) S. F. Blake, Bull. Torrey Bot. Club 51: 430 (1924).

Ditrichum macrophyllum Cass., Bull. Sci. Soc. Philom. Paris 1818: 59 (1818). Type: Je lai analyse dans lherbier de M. de Jussieu, o elle est tiguwt e avec doute, daprs Vahl, Conyza lobata, L. Holotype: P-JU.

Verbesina diversifolia DC., Prodr. 5: 615 (1836). Type/s?: è'' K circa Bahiam Brasiliae leg. cl. Blanchet [1652, 1799] et Salzmann [17]. ... (v.s.) Syntypes: G-DC. Isosyntypes: Salzmann 17, K 2.

Verbesina lancifolia Gardner, London J. Bot. 7: 406 (1848). Type: Hab. Bushy places near the city of Bahia. Sept. 1838. [Gardner] 875.

Bolivia (La Paz), Brazil.

[Section 11 Ochractinia B. L. Rob. & Greenm. Heads small; rays white or nearly so; leaves alternate.]

Verbesina mandonii Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea, 33(5): 528 (Feb. 1866), nom. nud. (based on Mandon 58) = ?Verbesina mandonii Sch.Bip. ex B. L. Rob. & Greenm. see note below, since Mandon 58 is clearly Flaveria bidentis (L.) Kuntze.

*Verbesina mandonii Sch.Bip. ex B. L. Rob. & Greenm., Proc. Amer. Acad. Arts 34: 547 (1899). Types: Andes of Bolivia, in woods, La Paz, altitude 2,600 to 3,700 m., January, 1861, G. Mandon, no. 57 in herb Gray; October, 1895, Rusby, no. 1721; 1889, Bang, no. 4. Syntypes: NY Bang 4 2, Mandon 57 3, Rusby 1721 3. Note: Schultz Bipontinus (1865: 79) listed Mandon 57 against Flaveria contrayerba.

Verbesina mandonii Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea, 33(5): 528 (Feb. 1866), nom. nud. (based on Mandon 58). Note: This may well be where Robinson & Greeman obtained their name, yet material of Mandon 58 is quite clearly of Flaveria bidentis (L.) Kuntze!

Bolivia (La Paz).

Note: The Mandon 57 syntypes in NY are of two differing taxa. NY 00274447 is Flaveria bidentis, as noted by Nee; the other two, NY 00274448, 0027449 are of Verbesina mandonii, sugesting that there may well have been an initial transposition of the numbers in Schultz Bipontinus list, or a mixing of materials when the duplicates were distributed.

[Section 8. Verbesinaria DC.]

Verbesina microptera (DC.) Herter, Rev. Sudam. Bot. 7: 235 (1843), comb. illegit. non Verbesina microptera DC. (1836) = Verbesina encelioides (Cav.) A. Gray

Verbesina nelidae Cabrera, Notas Mus. La Plata, Bot. 18(No. 86): 61 (1955). Type: Argentina. Jujuy: Valle Grande, 1500 m s.m., leg. A. Burkart et N. S. Troncoso, 11477, 26-II-1940. Holotype: LP; isotype: SI.

Argentina, Bolivia (?).

Verbesina nodiflora L., Cent. I. Pl. : 28 (1755) = Synedrella nodiflora (L.) Gaertn.

Verbesina octantha S. F. Blake, Amer. J. Bot. 12(10): 633 (1925), as nom. nov. pro Chaenophyllus macrophyllus Griseb. non Verbesina macrophylla (Cass.) S. F. Blake = Verbesina suncho (Griseb.) S. F. Blake

Verbesina oppositifolia Poir., Encycl. 8: 460 (1808) = Tilesia baccata (L.f.) Pruski

*Verbesina pflanzii Perkins, Bot. Jahrb. Syst. 49: 227 (1913). Type: Bolivien: Palca-La Paz, 3700 m . M. (K. Pflanz n. 47. Im April 1908 bl hend). Holotype: B.

Bolivia (La Paz).

[Sect. Verbesinaria]

Verbesina prostrata L., Sp. Pl.: 902 (1753) = Eclipta prostrata (L.) L.

*Verbesina rhomboidea J. Koster, Blumea 6: 270 (1948). Type: Hab.: kleiner Strauch auf dem Kamm der Laguna verde bei Comarapa, 2600 m alt., April 1911, [Herzog] n. 1984. Holotype: L.

Bolivia ().

Verbesina scabra Phil., Anales Mus. Nac. Chile, Secc. 2, Bot. 34: 186 (1870), non Benth. (1840) = Verbesina encelioides (Cav.) A. Gray

Verbesina scandens (L.) Klatt, Leopoldina 25: 106 (1889) = Salmea scandens (L.) DC.

*Verbesina semidecurrens Kuntze, Revis. Gen. Pl. 3(3): 183 (1898). Type: Bolivia: 3800 m Challapass zwischen oruro und Rio Tapacari. [Bolivia. Challapass zwischen Oruro und Rio Tapacari, 3800 m, 18 Mar 1892, Kuntze s.n. according to Wetter & Zanoni, 1985: 339]. Holotype: NY.

Bolivia (Oruro).

Verbesina soratae Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea 34(5): 528. (Feb. 1866), nom. nud. (based on Mandon 55) = Verbesina soratae Sch.Bip. ex B. L. Rob. & Greenm.

*Verbesina soratae Sch.Bip. ex B. L. Rob. & Greenm., Proc. Amer. Acad. Arts 34: 551 (1899). Types: Sorata, Bolivia, Mandon, no. 55, and in the same locality at 2,400 m. altitude, February, 1886, Dr. H. H. Rusby, no. 1722. Syntypes: ?; isosyntype (of Mandon 55): S.

Verbesina soratae Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea, 34(5): 528. (Feb. 1866), nom. nud. (based on Mandon 55).

Bolivia (La Paz).

[Sect. Saubinetia DC.]

*Verbesina subcordata DC., Prodr. 5: 614 (1836). Type: L circa Buenos-Ayres legit cl. Bacle [136]. ... (v.s. comm. cl. inv.). Holotype: G-DC.

Argentina, Bolivia (?).

Verbesina suncho (Griseb.) S. F. Blake, Amer. J. Bot., 12(10): 633 (1925)

Chaenocephalus suncho Griseb., Abh. K nigl. Ges. Wiss. Gttingen 24(1): 195 (March-April 1879); Symb. Fl. Argent. : 195 (1879). Type: Nom. vernac. Suncho v. Pino americano. C.: Oran, in sylvis virgineis Tabaccal ubi in locis caede arborum apricis vegetationem secundarium constituit.í¸.í¶' Blake (1925: 633) noted the type collection was in Y [= NY], sketch and fragm. G. [= GH]

Chaenocephalus heterophyllus Griseb., Abh. Knigl. Ges. Wiss. G ttingen 24(1): 196 (March-April 1879); Symb. Fl. Argent. : 196 (1879). Types: O.: Oran, pr. urbem; Tarija, Cuesta de Buyuyu. Syntypes: Lorentz & Hieronymus 343, 867, GOET. Blake (1925: 633) only cited Lorentz & Hieronymus 343 against Oran, but did not appear to have typified the name save citing (sketch and fragm. of TYPE; G) after this latter specimen. Blake mentioned the material from Tarija but not apparently the collector, except perhaps by reference to Kuntze. Kuntze (1898: 140) appears to only have referred to material from Tucuman, and nothing from Bolivia.

Chaenocephalus macrophyllus Griseb., Abh. Knigl. Ges. Wiss. G ttingen 24(1): 196 (March-April 1879); Symb. Fl. Argent.: 196 (1879). Types: T.: Tucuman, pr. urbem. Holotype: Lorentz & Hieronymus 193.

Verbesina octantha S. F. Blake, Amer. J. Bot. 12(10): 633 (1925), as nom. nov. pro Chaenophyllus macrophyllus Griseb. non Verbesina macrophylla (Cass.) S. F. Blake

Verbesina allophylla S. F. Blake, Amer. J. Bot. 12(10): 633 (1925), as nom. nov. pro. Chaenocephalus heterophyllus Griseb. non Verbesina heterophylla (Chapm.) A. Gray

Argentina, Bolivia (Tarija).

The following material was cited by Blake (1925: 633) under V. allophylla S. F. Blake:

Bolivia: Machichoirisa, alt. 1065 m., 1902, Williams 1592 (Y [= NY]); in cultivated ground, Huachi, head of Beni River, alt. 915 m., 1921, Rusby (Mulford Biol. Expl. 458; N [= US], Y [= NY]).

Vermifuga Ruiz & Pav., Fl. Peruv. Chil. Prodr. : 114 (1794) = Flaveria Juss.

Vermifuga corymbosa Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. : 216 (1798), based on Milleria contrayerba Cav. = Flaveria bidentis (L.) Kuntze

Vernonanthura H. Rob., Phytologia 73(2): 66 (1992) = Vernonia Schreb.

Vernonanthura amplexicaulis (R. E. Fr.) H. Rob., Phytologia 73(2): 68 (1992) = Vernonia amplexicaulis R. E. Fr.

Vernonanthura auriculata (Griseb.) H. Rob., Phytologia 73(2): 69 (1992) = Vernonia auriculata Griseb.

Vernonanthura brasiliana (L.) H. Rob., Phytologia 73(2): 69 (1992) = Vernonia brasiliana (L.) Druce

Vernonanthura canaminina (Gleason) H. Rob., Phytologia 73(2): 69 (1992) = Vernonia amplexicaulis R. E. Fr.

Vernonanthura ferruginea (Less.) H. Rob., Phytologia 73(2): 70 (1992) = Vernonia ferruginea Less.

Vernonanthura membranacea (Gardner) H. Rob., Phytologia 73(2): 71 (1992) = Vernonia membranacea Gardner

Vernonanthura patens (Kunth) H. Rob., Phytologia 73(2): 72 (1992) = Vernonia patens Kunth

Vernonanthura pinguis (Griseb.) H. Rob., Phytologia 73(2): 73 (1992) = Vernonia pinguis Griseb.

Vernonanthura prenanthoides (Gleason) H. Rob., Phytologia 73(2): 73 (1992) = Vernonia prenanthoides Gleason

Vernonanthura santacruzensis (Hieron.) H. Rob., Phytologia 76(1): 29 (1994) = Vernonia santacruzensis Hieron.

Vernonanthura squamulosa (Hook. & Arn.) H. Rob., Phytologia 73(2) :74 (1992) = Vernonia squamulosa Hook. & Arn.

Vernonanthura yurimaguasensis (Hieron.) H. Rob., Phytologia 73(2): 74 (1992) = Vernonia yurimaguasensis Hieron.

Vernonia Schreb. sect. Lepidaploa (Cass.) DC., Prodr. 5: 26 (1836) = Vernonia Schreb.

Vernonia Schreb. sect. Lepidaploa (Cass.) DC. subsect. Brevifoliae Cabrera, Darwiniana 6(3): 303 (1944) = Vernonia Schreb.

Vernonia Schreb. sect. Lepidaploa (Cass.) DC. subsect. Chamaedrys Cabrera, Darwiniana 6(3): 307 (1944) = Vernonia Schreb.

Vernonia Schreb. sect. Lepidaploa (Cass.) DC. subsect. Laurifoliae Cabrera, Darwiniana 6(3): 350 (1944) = Vernonia Schreb.

Vernonia Schreb. sect. Lepidaploa (Cass.) DC. subsect. Nitidulae Cabrera, Darwiniana 6(3): 347 (1944) = Vernonia Schreb.

Vernonia Schreb. sect. Lepidaploa (Cass.) DC. subsect. Nudiflorae Cabrera, Darwiniana 6(3): 353 (1944) = Vernonia Schreb.

Vernonia Schreb. sect. Lepidaploa subsect. Oligocephalae Baker in Mart., Fl. Bras. 6(2): 46 (1873), p.p. = Vernonia Schreb.

Vernonia Schreb. sect. Lepidaploa (Cass.) DC. subsect. Sellowianae Cabrera, Darwiniana 6(3): 306 (1944) = Vernonia Schreb.

Vernonia Schreb. sect. Stenocephalum (Sch.Bip.) Baker in Mart., Fl. Bras. 6(2): 25 (1873) = Vernonia Schreb.

Vernonia Schreb. sect. Tephrodes DC., Prodr. 5: 24 (1836) = Vernonia Schreb.

Vernonia Schreb. sect. Vanillosma Less., Linnaea 6: 630 (1831) = Piptocarpha R.Br.

Vernonia Schreb. sect. Vernonia subsect. Arborescentes Ekman, Ark. Bot. 13(15): 27 (1914) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Eremosis (DC.) S. B. Jones, Rhodora 81(No. 828): 435 (1979) = Critoniopsis Sch.Bip.

Vernonia Schreb. sect. Vernonia subsect. Noveboracenses Ekman, Ark. Bot. 13(15): 95 (1914) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Nudiflora Cabrera ser. Brevifoliae (Cabrera) S. B. Jones, Rhodora 81(No. 828): 438 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Nudiflorae Cabrera ser. Subulatae S. B. Jones, Rhodora 81(No. 828): 439 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Nudiflorae Cabrera ser. Verbascifoliae S. B. Jones, Rhodora 81(No. 828): 438 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Pallescentes Ekman, Ark. Bot. 13(15): 88 (1914) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Polyanthes Ekman, Ark. Bot. 13(15): 89 (1914) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Sagraeanae Ekman, Ark. Bot. 13(15): 11 (1914) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Scorpioides Ekman ser. Arborescentes (Ekman) S. B. Jones, Rhodora 81(No. 828): 445 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Scorpioides Ekman ser. Aureae S. B. Jones, Rhodora 81(No. 828): 442 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Scorpioides Ekman ser. Flexuosae (Cabrera) S. B. Jones, Rhodora 81(No. 828): 442 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Scorpioides Ekman ser. Foliatae (Benth. & Hook. f.) S. B. Jones, Rhodora 81(No. 828): 443 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Scorpioides Ekman ser. Pallescentes (Ekman) S. B. Jones, Rhodora 81(No. 828): 445 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Scorpioides (Benth. & Hook.f.) Ekman ser. Remotiflorae (Cabrera) S. B. Jones, Rhodora 81(No. 828): 441 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Scorpioides Ekman ser. Sagraeanae (Ekam) S. B. Jones, Rhodora 81(No. 828): 444 (1979) = Vernonia Schreb.

Vernonia Schreb. sect. Vernonia subsect. Stenocephalum (Sch.Bip.) S. B. Jones, Rhodora 81(No. 828): 437 (1979) = Vernonia Schreb.

Vernonia Schreb. subsect. Flexuosae Cabrera, Darwiniana 6: 329 (1944) = Vernonia Schreb.

Vernonia Schreb. subsect. Orbivestus S. B. Jones, Rhodora 83(No. 833): 61 (1981) = Vernonia Schreb.

Vernonia Schreb. subsect. Hilliardianae S. B. Jones, Rhodora 83(No. 833): 66 (1981) = Vernonia Schreb.

Vernonia Schreb. subsect. Tephrodes (DC.) S. B. Jones, Rhodora 83(No. 833): 70 (1981) = Vernonia Schreb.