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Schkuhria
Roth, Cat. Bot. 1: 116 (1797).
Tetracarpum
Moench, Meth. Suppl. : 240 (1802). Type: Schkuhria abrotanoides
Roth = Schkuhria pinnata (Lam.)
Kuntze
Chamaestephanum
Willd., Ges. Naturf. Freunde Berlin Mag. neuesten Entdeck. Gesammten Naturk. 1:
140 (1807). Type: not designated.
Mieria La Llave in La
Llave & Lex., Nov. Veg. Descr. 2: 12 (1825). Type: Mieria virgata Llave
& Lex. = Schkuhria virgata (Llave) DC. = Schkuhria pinnata (Lam.) Kuntze
Achyropappus Link
& Otto, Ic. Pl. Rar. Pl. 30 (1829), non Achyropappus
Kunth (1818). Type: Achyropappus
schkuhrioides Link & Otto = Schkuhria schkuhrioides (Link & Otto)
Thell.
Hopkirkia DC.,
Prodr. 5: 660 (1836), non Hopkirkia
Spreng. (1819). Type: Hopkirkia
anthemoidea DC. = Schkuhria anthemoidea (DC.) Coult.
Cephalobembix
Rydb., N. Amer. Fl. 34: 46 (1914). Type: Schkuhria neomexicana A. Gray
Type: Schkuhria
abrotanoides Roth = Schkuhria
pinnata (Lam.) Kuntze [var. abrotanoides (Roth) Cabrera]
References
Heiser Jr., C. B. (1945). A revision of the genus Schkuhria. Ann. Missouri Bot. Gard.
32(3): 265278.
Robinson, H. (2006). Schkuhria. In: G. Harling & L. Andersson (eds), Flora of
Ecuador, No. 77(1): 190(6). Compositae-Heliantheae, Part 1: Introduction,
genera MZ. Botanical Institute, Gteborg University, G teborg & Section
for Botany, Riksmuseum, Stockholm. pp. 9196.
Turner, B. L. (1995). Taxonomy and nomenclature of Schkuhria pinnata (Asteraceae,
Helenieae). Phytologia 79(5): 364368.
Note: Robinson (2006) took a very broad view of S. pinnata but noting varieties were
sometimes recognized. The full synonymy provided by Robinson is given below as S. pinnata, and could be divided.
Turners (1995) further recognition of the North American element as var. wislizeni was provided against the
background of MacVaughs (1984: 794799) treatment of plants from New Mexico.
All are recognized as synonyms of a broad species concept. Anyone wanting to
recognize any infraspecific categories should in the first instance examine
Heisers (1945) revision for the South American element; This, following Turner,
is to be referred solely to var. pinnata.
Key to species
1. Rays
absent; achenes 10 or more 2
Ray
or rays usually present; achenes 9 or less S. pinnata
2. (1) At least
upper half of pappus scales awned 3
Pappus
scales not awned, except rarely 1 or 24
3. (2) Awns of
pappus scales 1 mm or longer; scales colourless S. multiflora
Awns
less than 1 mm long; scales maroon to purplish S. degenerica
4. (2) Phyllaries
56, 2 rarely 3 mm wide; achenes usually less than 15; pappus scales rarely
longer than 0.5 mm S. multiflora [var. pusilla]
Phyllaries
7 or more, mostly 12 mm wide; achenes usually more than 15; pappus scales
frequently longer than 0.5 mm S. multiflora
Schkuhria
abrotanoides Roth, Catalecta Bot. 1: 116 (1797) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schuhria
abrotanoides Roth var. isopappa
(Benth.) Hieron., Bot. Jahrb. Syst. 29(1): 53 (1900) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schkuria
abrotanoides Roth var. pomasquiensis
Hieron., Bot. Jahrb. Syst. 29(1): 53 (1900) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schkuhria
advena Thell., Feddes Repert. Spec. Nov. Regni Veg. 11: 308 (1918) = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
anthemoidea (DC.) Coult. var. wislizenii
(A. Gray) Heiser f. flava (Rydb.)
Heiser, Ann. Missouri Bot. Gard. 32(3): 274 (1945) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schkuhria
anthemoidea (DC.) Coult. var. guatemalensis
(Rydb.) Heiser, Ann. Missouri Bot. Gard. 32(3): 273 (1945) = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
anthemoidea (DC.) Coult. var. wislizeni
(A. Gray) Heiser, Ann. Missouri Bot. Gard. 32(3): 273 (1945) = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
anthemoidea (DC.) Coult. var. wrightii
(A. Gray) Heiser, Ann. Missouri Bot. Gard. 32(3): 274 (1945) = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
bonariensis Hook. & Arn., J. Bot. (Hooker) 3(No. 22): 321 (1841), p.p.
= Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
coquimbana Phil., Anales Univ. Chile 90: 29 (1895) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
*Schkuhria
degenerica (Kuntze) R. E. Fr., Ark. Bot. 5(13): 22 (1906).
Schkuhria
pusilla Wedd. var. major
Sch.Bip., Bull. Soc. Bot. France 7: 80 (1865), nom. nud.
Schkuhria
oolepis Sch.Bip., Bull. Soc. Bot. France 12: 80 (1865); Linnaea 34(5): 529
(1866), nom. nud. pro syn. (based on Mandon
72).
Rothia degenerica
Kuntze, Revis. Gen. Pl. 3(3): 169 (1898). Type: Bolivia: 3000 m am Rio Tapacari.
[Bolivia. Rio Tapacari, 3000 m,
19 Mar 1892, Kuntze s.n. according
to Wetter & Zanoni, 1985: 336]
Argentina, Bolivia (Cochabamba, La Paz, Tarija).
Schkuhria
glabrescens Gand., Bull. Soc. Bot. France 65: 46 (1918) = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
guatemalensis (Rydb.) Standl. & Steyerm., Field Mus. Publ. Bot. 22: 319
(1940) = Schkuhria pinnata (Lam.)
Kuntze ex Thell.
Schkuhria
hopkirkia A. Gray, Smithsonian Contr. Knowl. [Pl. Wrightianae pt. 2] 5(Art.
6): 94 (1853) = Schkuhria pinnata
(Lam.) Kuntze ex Thell.
*Schkuhria
multiflora Hook. & Arn., J. Bot. 3: 322 (1841). Type: Achyropappus
schkuhrioides, Don, (non Link.) Mendoza; Dr
Gillies. Holotype: K.
Achyropappus
schkuhrioides D. Don ex Hook. & Arn., J. Bot. 3: 322 (1841), nom. nud.
pro syn., non A. schkurioides Link
& Otto (1829)(= Schkuhria schkuhrioides (Link & Otto) Thell.).
Schkuhria neo-mexicana
A. Gray, Mem. Amer. Acad. Arts, ser. 2, 4(1): 96 (1849). Type: Margin of
fields, Santa F; July, Aug. ([Fendler]
458.). Holotype: GH.
Schkuhria pusilla Wedd., Chloris Andina 1: 73, t. 14, B (1856). Types: íYTí²'Hab. Prou: dans les champs sablonneux, pr s du village de Moho (frontire de la Bolivie), au voisinage du lac de Titicaca, h. 3900 m tres. Bolivie: environs de Potosi (dOrbigny, no 1403).
Amblyopappus
neo-mexicana (A. Gray) A. Gray, Torrey Pacif. R. R. Report 4: 106 (1857).
Bahia neo-mexicana
(A. Gray) A. Gray, Proc. Amer. Acad. Arts 19: 27 (1883).
Rothia intermedia
Kuntze, Revis. Gen. Pl. 3(3): 170 (1898). Type: Bolivia: 4000 m Challapata.
[Bolivia. Challapata, 4000 m, 11
Mar 1892, Kuntze s.n. according to
Wetter & Zanoni, 1985: 336]
Rothia pusilla (Wedd.) Kuntze, Revis. Gen. Pl. 3(3): 170 (1898).
Schkuhria pusilla
Wedd. var. aristata R. E. Fr., Nova
Acta Regiae Soc. Sci. Upsal. 4(11): 85, t. 6, 8 (1905). Type: [Argentina:]
Moreno loco arenosis, ca. 3500 m. s. m. (12 Dec. 1901; Fr[
Achyropappus
neo-mexicana (A. Gray) Rydb., Fl. Colorado : 377 (1906).
Cephalobembix
neo-mexicana (A. Gray) Rydb., N. Amer. Fl. 34: 46 (1914).
Schkuhria pusilla
Wedd. var. longepedicellata Hauman,
Anal. Soc. Ci. Argent. 86: 328 (1918) [p. 174 in separately reprint see
References]. Types: En terrains secs, assez rare, prs de Puente del Inca, dans
le sable au fond de la vall e, janvier 1910 (examplaires rachitique peu
ramifis); Pr cordillre de San Juan, Carmen Alto, leg. Bodenbender, f vrier 1897 (Herb. Fac. Med. Buenos Aires).
Holotype: ?BAF.
*Schkuhria
multiflora Hook. & Arn. var. pusilla
(Wedd.) Cabrera, Anales Soc. Cient. Argentina 114: 192 (1932).
Schkuhria multiflora
Hook. & Arn. var. aristata (R. E.
Fr.) Cabrera, Anal. Soc. Cient. Argent. 114: 193 (1932).
Argentina, Bolivia (Chuquisaca, La Paz, Oruro, Tarija), Chile, Mexico, Peru, USA.
Canchalagua
(Ct) (Petenatti & Ariza Espinar,
1997: 23).
Schkuhria
multiflora Hook. & Arn. var. aristata
(R. E. Fr.) Cabrera, Anal. Soc. Cient. Argent. 114: 193 (1932) = Schkuhria multiflora Hook. & Arn.
*Schkuhria multiflora Hook. & Arn.
var. pusilla (Wedd.) Cabrera, Anales
Soc. Cient. Argentina 114: 192 (1932) = Schkuhria multiflora Hook. & Arn.
Schkuhria
octoaristata DC., Prodr. 5: 654 (1836) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schkuhria oolepis Sch.Bip., Bull. Soc. Bot. France 12: 80 (1865); Linnaea 34(5): 529 (Feb. 1866), nom. nud. pro syn. (based on Mandon 72) = Schkuhria degenerica (Kuntze) R. E. Fr.
Schkuhria
pinnata (Lam.) Kuntze, Revis. Gen. Pl. 3(3): 170 (1898), pro syn. = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
*Schkuhria pinnata
(Lam.) Kuntze ex Thell., Feddes Repert. Spec. Nov. Regni Veg. 11: 308 (1912).
Pectis pinnata
Lam., J. Hist. Nat. 2: 150, pl. 31 (1792). Type: Habitat . . . . K s. L. Holotype: ?Z.
Schkuhria abrotanoides
Roth, Catal. Bot. 1: 116 (1797). Type: Sub nomine Bellii minuti mihi communicata sunt semina ab amico perdilecto Ill.
de Voigt. Holotype: unknown, but
probably B or B-W.
Mieria
virgata La Llave & Lex., Nov. Veg. Descr., fasc. 2: 9 (1825).
Type: Habitat ad colles Tacnbaya et aliis in locis Mexico vicinis. Holotype:
G.
Schkuhria octoaristata
DC., Prodr. 5: 654 (1836). Type:
in Peruvi legit cl. Dombey [27]. ...
(v.s.). Holotype: G-DC. There are two specimens labelled as S. octoaristata in G-DC. One is clearly
a Dombey collection from Paris, which
is considered to be the holotype; the other, numbered 84, is without the
collectorâ''s name.
Schkuhria
virgata (Llave & Lex.) DC., Prodr. 5: 654 (1836). [Note: this
was placed in synonymy with S.
octoaristata in Robinsons homotypic synonymy but this cannot be the case!]
Hopkirkia anthemoidea
DC., Prodr. 5: 660 (1836). Type: in Mexici agro Regiomontano legit cl. Haenke. (v.s. in h. Haenk. cl. de
Sternberg comm.). Holotype: PR; isotype: G-DC.
Schkuhria bonariensis
Hook. & Arn., J. Bot. (Hooker) 3(No. 22): 321 (1841), pro parte. Types: Ò'Pampas
of Buenos Ayres; Dr. Gillies. Buenos
Ayres; Tweedie.
Schkuhria isopappa
Benth., Pl. Hartweg. : 205 (1845). Type: [Peru:] Ad pontem Guapulo prope
Quito. [Hartweg ].
Schkuhria
wislizeni A. Gray, Mem. Amer. Acad. Arts [Pl. Fendl.] ser. 2, 4: 96 (1849).
Type: High mountains around Cosiquiriachi in the Sierra Madre west of
Chihuahua, Oct. Dr. Wislizenus.
Schkuhria hopkirkia
A. Gray, Smithsonian Contr. Knowl. [Pl. Wrightianae pt. 2] 5 (Art. 6): 94
(1853), nom. illegit., based on Hopkirkia
anthemoidea DC.
Schkuhria wrightii A.
Gray, Smithsonian Contr. Knowl. [Pl. Wrightianae pt. 2] 5 (Art. 6): 95 (1853).
Type: On the Sonorita near Deserted Rancho, Sonora; Sept. ([Wright] 1254.).
Amblyopappus mendocinus
Phil., Anales Univ. Chile 36: 184 (1870). Type: [Chile:] Mendoza. Pizarro
(1960: 131) cited two collections in SGO 60578, 44110. Note: Philippi (1870)
also noted Nuestros ejemplares no son ... suggesting there were either two
collections or more than one specimen per collection.
Schkuhria pringlei S.
Watson, Proc. Amer. Acad. Arts 23: 278 (1888). Type: In moist places on the
plains near Guerrero, Chihuahua; C. G.
Pringle (n. 1292), Sept., 1887.
Schkuhria
coquimbana Phil., Anales Univ. Chile 90: 29 (1895). Type: Prope
Paihuano in provincia Coquimbo legit Frid.
Philippi incolae eam cachanlahuen
cimarron vocant. [Cerca de Paihuano, en la provincia de Coquimbo, la
recolect Federico Philippi; la gente la llama cachanlahu n cimarrn. 60580
ó"'- Pizarro, 1960: 154]
Rothia
pinnata (Lam.) Kuntze, Revis. Gen. Pl. 3(3): 170 (1898).
Schkuhria pinnata
(Lam.) Kuntze, Revis. Gen. Pl. 3(3): 170 (1898), pro syn.
Rothia pinnata
(Lam.) Kuntze a
pallida Kuntze, Revis. Gen. Pl. 3(3):
170 (1898), pro parte
Rothia pinnata
(Lam.) Kuntze b
purpuascens Kuntze, Revis. Gen. Pl. 3(3):
170 (1898), pro parte
Schkuria
abrotanoides Roth var. pomasquiensis
Hieron., Bot. Jahrb. Syst. 29(1): 53 (1900*). Type: [Ecuador]
Crescit in arvis arenosis prope Pomasqui (S.[odiro] n. 47). Holotype: B. [*Note:
See Reference section concerning problem with date of publication]
Schuhria abrotanoides Roth var. isopappa (Benth.) Hieron., Bot. Jahrb.
Syst. 29(1): 53 (1900*). [*Note: See Reference section concerning problem with
date of publication]
Tetracarpum
wrightii (A. Gray) Rydb., N. Amer. Fl. 34: 44 (1914).
Tetracarpum
pringlei (S. Watson) Rydb., N. Amer. Fl. 34: 44 (1914).
Tetracarpum
virgatum (La Llave & Lex.) Rydb., N. Amer. Fl. 34: 45 (1914).
Tetracarpum
anthemoideum (DC.) Rydb., N. Amer. Fl. 34: 45 (1914).
Tetracarpum
wislizeni (A. Gray) Rydb., N. Amer. Fl. 34: 45 (1914).
Tetracarpum
guatemalensis Rydb., N. Amer. Fl. 34: 45 (1914). Type: Type
collected at Llano de San Juan de Dios, Guatemala, 1865, Bernouli 135. Holotype: originally LGO, transferred to USNC.
Tetracarpum flavum
Rydb., N. Amer. Fl. 34: 46 (1914). Types: Type collected in Sierra de San
Felipe, Oaxaca,
Schkuhria
glabrescens Gand., Bull. Soc. Bot. France 65: 46 (1918). Type: Mexico,
Valley of Mexico, alt. 7300 ped. (Pringle
n. 7928!).
Schkuhria advena
Thell., Feddes Repert. Spec. Nov. Regni Veg. 11: 308 (1918). Type: Sd-Afrika:
Transvaal: Florida, 1906, H. Hutton
no. 630 (Albany Museum), ab S.
bonariensis in herb. Univ. Z rich. Holotype: Z.
Schkuhria guatemalensis
(Rydb.) Standl. & Steyerm., Field Mus. Publ. Bot. 22: 319 (1940).
Schkuhria pinnata (Lam.) Kuntze ex
Thell. var. virgata (Llave &
Lex.) Heiser, Ann. Missouri Bot. Gard., 32(3): 271 (1945).
Schkuhria
pinnata (Lam.) Kuntze ex Thell. f. pringlei
(S. Watson) Heiser, Ann. Missouri Bot. Gard. 32(3): 271 (1945).
Schkuhria
anthemoidea (DC.) Coult. var. guatemalensis
(Rydb.) Heiser, Ann. Missouri Bot. Gard. 32(3): 273 (1945).
Schkuhria
anthemoidea (DC.) Coult. var. wislizenii
(A. Gray) Heiser, Ann. Missouri Bot. Gard. 32(3): 273 (1945).
Schkuhria
anthemoidea (DC.) Coult. var. wislizenii
(A. Gray) Heiser f. flava (Rydb.)
Heiser, Ann. Missouri Bot. Gard. 32(3): 274 (1945).
Schkuhria
anthemoidea (DC.) Coult. var. wrightii
(A. Gray) Heiser, Ann. Missouri Bot. Gard. 32(3): 274 (1945).
Schkuhria wislizeni
A. Gray var. wrightii (A. Gray) S. F.
Blake, Leaflets West. Bot. 6: 115 (1951).
Schkuhria
wislizeni (A. Gray) Rydb. var. frustrata
S. F. Blake, Leaflets West. Bot. 6: 115 (1951), nom. nov. pro Hopkirkia anthemoidea DC.
Schkuhria wilsizeni
A. Gray var. guatemalensis (Rydb.) S.
F. Blake, Leaflets West. Bot. 6: 115 (1951).
Schkuhria wislizeni
A. Gray f. flava (Rydb.) S. F. Blake,
Leaflets West. Bot. 6: 115 (1951).
Schkuhria pinnata
(Lam.) Kuntze ex Thell. var. guatemalensis
(Rydb.) McVaugh, Contr. Univ. Mich. Herb. 9: 443 (1972).
Schkuhria
pinnata (Lam.) Kuntze var. wislizeni
(A. Gray) B. L. Turner, Phytologia 79: 364 (1995).
Argentina, Bolivia (Chuquisaca, Cochabamba, La Paz, Santa
Cruz, Tarija), Brazil, Chile, Colombia, Ecuador, Peru, Uruguay, USA, Venezuela;
Cape Verde Islands, cult. ex France, Mozambique.
Canchalagua;
Matapulgas (Petenatti & Ariza Espinar, 1997: 24)
Schkuhria
pinnata (Lam.) Kuntze ex Thell. var. guatemalensis
(Rydb.) McVaugh, Contr. Univ. Mich. Herb. 9: 443 (1972) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schkuhria
pinnata (Lam.) Kuntze ex Thell. f. pringlei
(S. Watson) Heiser, Ann. Missouri Bot. Gard. 32: 271 (1945) = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
pinnata (Lam.) Kuntze ex Thell. var. virgata
(Llave & Lex.) Heiser, Ann. Missouri Bot. Gard., 32: 271 (1945) = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
pinnata (Lam.) Kuntze var. wislizeni
(A. Gray) B. L. Turner, Phytologia 79: 364 (1995) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schkuhria
pusilla Wedd., Chloris Andina 1: 73, t. 14, B (1856) = Schkuhria multiflora Hook. & Arn.
Schkuhria
pusilla Wedd. var. aristata R. E.
Fr., Nova Acta Regiae Soc. Sci. Upsal. 4(11): 85, t. 6, 8 (1905) = Schkuhria multiflora Hook. & Arn.
Schkuhria
pusilla Wedd. var. longepedicellata
Hauman, Anal. Soc. Ci. Argent. 86: 328 (1918) = Schkuhria multiflora Hook. & Arn.
Schkuhria pusilla Wedd. var. major Sch.Bip., Bull. Soc. Bot. France 7: 80 (1865), nom. nud. = Schkuhria degenerica (Kuntze) R. E. Fr.
Schkuhria
virgata (La Llave & Lex.) DC., Prodr. 5: 654 (1836) = Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
wislizeni A. Gray, Mem. Amer. Acad. Arts [Pl. Fendl.] ser. 2, 4: 96 (1849) =
Schkuhria pinnata (Lam.) Kuntze ex
Thell.
Schkuhria
wislizeni A. Gray f. flava
(Rydb.) S. F. Blake, Leaflets West. Bot. 6: 115 (1951) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schkuhria wislizeni (A. Gray) Rydb. var. frustrata S. F. Blake, Leaflets West. Bot. 6: 115 (1951) = Schkuhria pinnata (Lam.) Kuntze ex Thell.
Schkuhria
wrightii A. Gray, Smithsonian Contr. Knowl. [Pl. Wrightianae pt. 2] 5 (Art.
6): 95 (1853) = Schkuhria pinnata
(Lam.) Kuntze ex Thell.
Senecio L. sect. Culcitium (Humb. & Bonpl.) Cuatrec., Fieldiana, Bot. 27: 50 (1950) = Culcitium Humb. & Bonpl.
Senecio
L. sect. Dendrophorbium Cuatrec.,
Fieldiana, Bot. 27(2): 72 (1951) = Dendrophorbium
(Cuatrec.) C. Jeffrey
Senecio
L. sect. Diclini Cabrera, Lilloa 5(1):
73 (1939) = Chersodoma Phil.
Senecio L. subgen. Dioicosenecio Cabrera, Lilloa 5(1): 68 (1939), nom. nud. = Chersodoma Phil.
Senecio L. sect. Emilioidei Muschl., Bot. Jahrb. Syst. 43: 40 (1909) = Emilia (Cass.) Cass.
Senecio L. sect. Macbrideus Cuatrec., Fieldiana, Bot. 27(2): 72 (1951) = Dendrophorbium (Cuatrec.) C. Jeffrey
Senecio L. sect. Myriocephalus Cabrera, Lilloa 15: 56 (1949) = Dendrophorbium (Cuatrec.) C. Jeffrey
Senecio L. ser. Myriocephali (Cabrera) Cabrera, Darwiniana 26(14): 126 (1985), comb. illegit, nom. non rite public. = Dendrophorbium (Cuatrec.) C. Jeffrey
Senecio
L. sect. Pluricephali Cabrera, Lilloa
5(1): 70 (1939) = Dendrophorbium
(Cuatrec.) C. Jeffrey
Senecio L. sect. Praegynoxys Cuatrec., Fieldiana, Bot. 27(2): 72 (1951) = Nordenstamia Lundin
Senecio
L. sect. Streptothamnus Greenm., Bot.
Jahrb. Syst. 32: 19 (1902) = Pentacalia
Cass.
