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Pentacalia
Cass., Dict. Sci. Nat. 48: 461 (1827).
Senecio L. sect. Streptothamnus Greenm., Bot. Jahrb.
Syst. 32: 19 (1902).
Type: Senecio
streptothamnus Greenm. ex Standl. = Pentacalia streptothamna (Greenm. ex
Standl.) H. Rob. & Cuatrec.
References
Cabrera, A. L. (1954). Senecios Sudamericanas nuevos o
crticos. Darwiniana 10(4): 547605 & L mina I
Cabrera, A. L. (1985). El genero Senecio (Compositae) en Bolivia. Darwiniana 26(14): 79217. [q.v. sect. Streptothamni Greenm.]
Robinson, H. & J. Cuatrecasas (1993). New species
of Pentacalia (Senecioneae:
Asteraceae) from Ecuador, Peru, and Bolivia. Novon 3(3): 284301.
Vision, T. J. & M. O. Dillon (1996). Sinopsis de Senecio L. (Senecioneae, Asteraceae)
para el Per. Arnaldoa 4(1): 2346.
Key to species
1. Capitula
radiate; margins florets with well-developed ray limbs 2
Captula
discoid or sub-discoid; marginal florets tubular or absent; involucre 46 mm
tall 14
2. (1) Involucres
710 mm tall 3
Involucres
3æ--6 mm tall; leaves never with revolute margins 6
3. (2) Leaves
ovate or elliptic, with repand-dentate or entire margins, more or less
conspicuously revolute; phyllaries 8144
Leaves
oblong-lanceolate, base attenuate or rounded, entire; phyllaries 8 Senecio
buchtienii
4. (3) Leaves
911 cm long by 34 cm wide, tomentose beneath; inflorescences paniculate;
phyllaries 14 P. cardenasii
Leaves
28 cm long by 1.54 cm wide, glabrous above and beneath 5
5. (4) Leaves
ovate or ovate-oblong, usually with a cordate base, margins crennate-dentate;
phyllaries 8 (13)P. dictyophlebius
Leaves
ovate-elliptic, base attenuate or rounded, margins entire; phyllaries 13P.
tablensis
6. (2) Leaf
margins serrate; phyllaries 137
Leaf
margins entire 9
7. (6) Petioles
22.5 cm long Senecio croicensis
Petioles
110 mm long 8
8. (7) Leaf
bases rounded; pedicels 1525 mm long; inflorescence of lax panicles P.
yungasensis
Leaf
bases cuneate; pedicels 515 mm long; inflorescence of dense panicles P.
chulumanicus
9. (6) Ray
limbs very short or absent, not exceeding disc florets; leaves lanceolate P. jelskii
Ray
limbs well-developed, exceeding disc florets 10
10. (9) Involucre
6 mm tall 11
Involucre
3.54 mm tall12
11. (10) Stems more
or less floccose; involucre 6 mm diam. P. floccosa
Stems
not floccose; involucre 4 mm diam. P. epiphytica
12. (10) Leaves
elliptic-lanceolate, 611 cm long by 2.54 cm wide; involucres 3.54 mm tall P.
miguelii
Leaves
ovate or elliptic; involucre 3 mm tall 13
13. (12) Phyllaries
89; leaves elliptic or ovate, 89 cm long by 46 cm wide P. psidiifolia
Phyllaries
1113; leves ovate, 59 cm long by 2.54.5 cm wide P. brittoniana
14. (1) Leaves
densely tomentose beneath 15
Leaves
somewhat lanuginose or glabrous beneath 17
15. (14) Leaf
margins entire P. marinii
Leaf
margins dentate 16
16. (15) Involucre
glabrescent; leaves white-tomentose beneath P. herzogii
Involucre
and underside of leaves yellowish-brown tomentose P. oronocensis
17. (14) Leaf
margins shortly denticulate 18
Leaf
margins entire 19
18. (17) Leaves lanceolate,
23 cm wide, reticulate venation conspicuous/prominent beneath P.
beckii
Leaves
elliptic, 3.55 cm wide, venation inconspicuous beneath P.
sailapatensis
19. (17) Leaves
lanceolate; involucre 5 mm tall P. jelskii
Leaves
ovate20
20. (19) Involucre 4
mm tall; leaves 3.56 cm wide P. comarapensis
Involucre
56 mm tall; leaves 1.53 cm wide P. subglomerosa
Pentacalia beckii
(Cabrera) Cuatrec., Phytologia 69(5): 314 (1990).
Senecio beckii
Cabrera, Hickenia 2(4): 15 (1984). Type: Bolivia. Deptro. La Paz, Prov. Nor
Yungas, La Paz 65 km hacia Coroico. 2700 m s.m. Bosque temporalmente nebuloso
en zona de nubes, leg. St. G. Beck No
1822, 28-
Bolivia (La Paz).
Pentacalia
brittoniana (Hieron.) Cuatrec., Phytologia 49(3): 244 (1981).
*Senecio sprucei Britton, Bull. Torrey Bot. Club 19(9):
265 (1892), non Klatt (1888). [Types: Yungas, 4,000 ft. ([Rusby] 1695). The same as Spruces No. 4811.
*Senecio brittonianus
Hieron., Bot. Jahrb. Syst. 29(1): 72 (1900), as nom. nov. pro S. sprucei Britton.
Bolivia (La Paz).
Pentacalia
buchtienii (Greenm,) Cuatrec., Phytologia 49(3): 244 (1981) = Denrophorbium buchtienii (Greenm.) C.
Jeffrey
Pentacalia
cardenasii (Cuatrec.) Cuatrec., Phytologia 49(3): 253 (1981).
*Senecio cardenasii
Cuatrec., Fieldiana, Bot. 27(2): 48 (1951).
Type: Bolivia, near Chulumani way to Yungas of Tablas, Cochabamba, 2700 m.
alt. on wet bushy slopes. Herb 2 met. flowers yellow. Collect. M. Cardenas 3988. Syntypes: F, US.
Bolivia (Cochabamba).
Pentacalia
chulumanica (Cabrera) Cuatrec., Phytologia 69(5): 314 (1990).
Senecio chulumanicus
Cabrera, Hickenia 2(4): 17 (1984). Type: BOLIVIA. Depto. La Paz, Prov. Sud
Yungas, Chulumani ca. 5 km hacia Irupana, entrando por la propriedad del Sr.
Portugal. 1850 m s.m. Leg. St. G. Beck
No 4875, 3-X-1981. Holotype: SI; isotype: LPB. Note: Freire & Iharlegui
(2000: 336) noted that on the collecting label the date was clearly 30-X-1981.
Bolivia (La Paz).
Pentacalia
comarapensis (Cabrera) Cuatrec., Phytologia 49(3): 245 (1981).
*Senecio comarapensis
Cabrera, Darwiniana 10(4): 599 (1954). Type:
Bolivia: Dep. Santa Cruz,
Comarapa, Yungas de San Mateo, 2900 m s.m., leg. J. Steinbach, 8519, 25-X-1938. Syntypes: LIL, S, MO. Lectotype
(selected by Cabrera, 1985: 106, as holotype): LIL (91764); isolectotypes:
MO, S.
Bolivia (Santa Cruz).
Pentacalia
dictyophlebia (Greenm.) Cuatrec., Phytologia 49(3): 245 (1981).
*Senecio dictyophlebius
Greenm., Ann. Missouri Bot. Gard. 25(4): 801 (1938). Type: Bolivia: vicinity of Sorata, alt. 3400
m., 31 Dec., 1858, G. Mandon, no.
146. Holotype: K; isotypes: BM, BR, G, NY, W.
Bolivia (La Paz).
Pentacalia
epiphytica (Kuntze) Cuatrec., Phytologia 49(3): 246 (1981).
*Senecio epiphyticus
Kuntze, Revis. Gen. Pl. 3(3): 173 (1898). Type: Bolivia: 3000 m Tunarigebirge,
Cinchona-Region, hoch in den Baumkronen nur wachsend. [Bolivia.
Tunarigebirge, Cinchona-Region, 3000 m, May 1892, Kuntze s.n. according to Wetter & Zanoni, 1985: 337].
holotype: NY.
*Senecio steinbachianus
Cuatrec., Fieldiana, Bot. 27(2): 55 (1951). Type: Bolivia, Prov. Chapare,
Dept. Cochabamba: Quebrada de Corani, 200 met. lat. Wurzelt und rankt an
Staemen. Coll. 21-VI-1929 Jose Steinbach
9876. Holotype: F; isotype: S.
Argentina, Bolivia (Cochabamba).
Cabrera (1975: 258) also recorded this species from Peru, but
later (1985: 96) only recorded it for Argentina and Bolivia.
Pentacalia
floccosa (Britton) Cuatrec., Phytologia 49(3): 246 (1981).
*Senecio floscosus
Britton, Bull. Torrey Bot. Club 19(9): 264 (1892).
Types: Near La Paz, 10,000 ft. ([Rusby]
1680); Unduavi, 8,000 ft. ([Rusby]
1720).
Bolivia (Cochabamba, La Paz), Peru.
Pentacalia herzogii (Cabrera) Cuatrec., Phytologia 49(3): 246 (1981).
Senecio herzogii
Beauverd ex Herzog, Pflanzenw. Bolivischen Anden : 188 (1923), nom. nud.
*Senecio herzogii
Cabrera, Blumea 7(1): 202 (1952). Type: [Bolivia]
Hab.: Kletternd im Bergwald bei Choquetanga Grande, 3300 m alt., Okt. 1911, [Herzog] n. 2145 (Typus). Holotype: L.
Bolivia (?).
Pentacalia
inquisiviensis H. Rob. & Cuatrec., Novon 3(3): 300 (1993). Type:
Bolivia. La Paz: Prov. Inquisivi, on slopes W of Ro Glorieta, and along river
between mouths of R o Aballahuanta and Ro Cayani 7 km S of Choquetanga,
1654'S 67í°°17'W, 3,150 3,550 m, 19 June 1991, Lewis 38950. Holotype: US; isotype: MO.
Bolivia (La Paz).
Pentacalia
krukoffii (Cuatrec.) Cuatrec., Phytologia 49(3): 255 (1981) = Dendrophorbium krukoffii (Cuatrec.) C.
Jeffrey
Pentacalia
jelskii (Hieron.) Cuatrec., Phytologia 49(3): 247 (1981).
Senecio jelskii
Hieron., Bot. Jahrb. Syst. 36(5): 509 (1905).
Type: Peruvia: crescit prope Tambillo (J.[elski] n. 769, anno 1878). Holotype: B.
*Senecio prunioides
Rusby, Bull. New York Bot. Gard. 4(14): 396 (1907). Type: [Bolivia:]
Climbing 10 to 15 feet upon trees, in forest-mould and clay, the flowers of a
dirty yellow, Coroico, Sept. 1894. ([Bang]
No. 2437.) Holotype: NY; isotype: US.
Bolivia (La Paz, Santa Cruz), Peru.
Pentacalia
lewisii H. Rob. & Cuatrec., Novon 3(3): 301 (1993). Type: Bolivia. La
Paz: Prov. Inquisivi, Pavionani, on slope above Pavionani fork of Ro Chimu
below headwater divide with R o Janko Kalani, area 7 km N of Choquetanga,
1648'S 6718'W, 3,700 3,750 m, 9 Apr. 1991, Lewis 38520. Holotype: US; isotype: MO.
Bolivia (La Paz).
Pentacalia
marinii (Cabrera) Cuatrec., Phytologia 49(3): 248 (1981). Note: although
not citing an exact basionym page, the page range included the whole of
Cabreras protologue (pp. 590593) to Senecio
marinii. This is perfectly acceptable under the present Code.
*Gynoxys discolor
Rusby, Bull. New York Bot. Gard. 4(14): 398 (1907). Type: [Bolivia:] ([Bang] No. 2280.) Holotype: NY; isotype:
US.
Senecio marinii
Cabrera, Darwiniana 10: 590 (1954). Type: Per. Dep. Cuzco, Prov. Convenci n,
Amaybamba, 1800 m s.m., leg. F. Marn 1789,
X-1949. Holotype: LP. Note: in his observations Cabrera said that he would
í''probably have to refer Gynoxys discolor
Rusby to this species, but noted that the florets in one isotype were too
poorly developed to allow proper comparison.
Bolivia (?), Peru.
Pentacalia
medullosa (Sch.Bip. ex Greenm.) Cuatrec., Phytologia 49(3): 256 (1981) = Dendrophorbium medullosum (Sch.Bip. ex
Greenm.) C. Jeffrey
Pentacalia
miguelii (Cuatrec.) Cuatrec., Phytologia 49(3): 248 (1981).
*Senecio miguelii
Cuatrec., Fieldiana, Bot. 27(2): 54 (1951). Type: Bolivia, collected Miguel Bang 2039. Syntypes: F, US. [2
duplicates in US marked as isotype]; isotype: LD.
Bolivia (?La Paz).
Pentacalia
oronocensis (DC.) Cuatrec., Phytologia 49(3): 248 (1981).
*Senecio oronocensis
DC., Prodr. 6: 423 (1838). Type: L in Amer. australis montibus Oronocensibus
legit cl. Haenke. ... (v.s. in h.
Haenke cl. de Sternberg miss.). Holotype: PR.
*Senecio baccharidiflorus
Rusby, Bull. New York Bot. Gard. 4(14): 397 (1907). Type: [Bolivia:]
Ò' A climber with white flowers, growing in rich forest-mould. Unduavi,
September, 1894. ([Bang] No. 2494.)
Senecio cuzcoensis
Cabrera, Notas Mus. La Plata, Bot. 9(No. 45): 196
(1941). Type: Per.
Departamento de Cuzco, Provincia de Pancastambo: Pillahuata, 2800 m. s. m.,
voluble en bosques ralos, leg. C. Vargas,
no 1908, 18-VI-1940. Holotype: Herb. Cabreraâ'' LP (64716).
Senecio ramonii Cuatrec.,
Fieldiana, Bot. 27(2): 53 (1951). Type: Peru: Carpish, entre Huanuco y Tingo
Mara, 2800-2900 m. alt. Flores blancas. Arbusto 1-1.5 met. Collect. Aug. 1947 Ram n Ferreyra 2310. Holotype: US.
Bolivia (La Paz), Peru.
Pentacalia
psidiifolia (Rusby) Cuatrec., Phytologia 49(3): 249 (1981).
*Senecio psidiifolius
Rusby, Mem. Torrey Bot. Club 6(1): 66 (1896). Type: [Bolivia:] Mapiri,
JulyAug., 1892 ([Bang] 1532).
Holotype: NY; isotypes: US 2, Z (000003971).
Bolivia (La Paz).
Pentacalia
sailapatensis (Cuatrec.) Cuatrec., Phytologia 49(3): 249 (1981).
*Senecio sailapatensis
Cuatrec., Fieldiana, Bot. 27(2): 52 (1951). Type: Bolivia, Dept. Cochabamba,
Prov. Ayopaya: Sailapata 2700 m. alt. Vine flowers white, collect. X-1935 in
wet forest by M. Crdenas 3273.
Holotype: US.
Bolivia (Cochabamba).
Pentacalia
subglomerosa (Greenm.) Cuatrec., Phytologia 49(3): 250 (1981).
*Senecio subglomerosus
Greenm., Ann. Missouri Bot. Gard. 10(1): 93 (1923). Type: á''Bolivia: Coroico,
Yungas, September, 1894, Bang 2459.
Holotype: NY; isotypes: GH, MO, US 2.
Bolivia (La Paz).
Pentacalia
tablensis (Cabrera) Cuatrec., Phytologia 49(3): 250 (1981).
*Senecio tablensis
Cabrera, Blumea 7(1): 203 (1950). Type: Hab.: Liane and der Waldgrenze ber
Tablas, 3400 m alt., Mai 1911, [Herzog]
n. 2204. Holotype: L; isotype: LP (899000).
Bolivia (Cochabamba).
Pentacalia
yungasensis (Britton) Cuatrec., Phytologia 69(5): 314 (1990).
*Senecio yungasensis
Britton, Bull. Torrey Bot. Club 19: 264 (1892). Type: Yungas, 4,000 ft. ([Rusby] 1719). Holotype: NY; isotype:
US.
Bolivia (La Paz), Peru.
