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Heterosperma Cav., Ic. Pl. 3: 34 (1794).

Heterospermum Willd., Sp. Pl. 3: 2129 (1803), orth. var.

Microdonta Nutt., Trans. Amer. Phil. Soc., ser. 2, 7: 369 (1841) Type: Microdonta nana Nutt. = Heterosperma nanum (Nutt.) Sherff

Type: Heterosperma pinnatum Cav. [published as pinnata but since Heterosperma is considered neuter the epithet spelling is corrected]

References

Blake, S. F. (1915). Four new heterospermas. J. Bot. 53(No. 635): 322324.

Cabrera, A. L. (1978). Heterosperma Cav. In: A. L. Cabrera, Flora de la Provincia de Jujuy, Republica Argentina. Parte X Compositae. pp. 399404.

Robinson, H. (2006). Heterosperma. In: G. Harling & L. Andersson (eds), Flora of Ecuador, No. 77(1): 190(6). Compositae-Heliantheae, Part 1: Introduction, genera AL. Botanical Institute, Gteborg University, G teborg & Section for Botany, Riksmuseum, Stockholm. pp. 186189.

Note: Robinson (2006) mentioned H. obtusifolium Cav. although I can find no record of this name in Cabrera, or Index Kewensis nor International Plant Name Index. It is probable he was referring to H. ovatifolium Cav.

Heterosperma depressum Griseb., Abh. Knigl. Ges. Wiss. G ttingen 19([1]): 187 (1874); Pl. Lorentz.: 139 (1874) = Heterosperma diversifolium Kunth

*Heterosperma diversifolium Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 193 (1818). Type: Crescit in aridis, prope urbem Quito et in convalli Guallagambensi, alt. 1150 1500 hex. 0 Floret Januario, Februario. Holotype: P-Bonpl.

Heterosperma depressum Griseb., Abh. Knigl. Ges. Wiss. G ttingen 19([1]): 187 (1874); Pl. Lorentz.: 139 (1874). Type: Tucuman, formationem alpinam herbaceam pr. Cienega ex magna parte constituens. Holotype: Lorentz 135, GOET.

Heterosperma pinnatum Cav. var. d depressum (Griseb.) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898).

Heterosperma pinnatum Cav. var. e diversifolium (Kunth) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898).

Heterosperma spathulatum S. F. Blake, J. Bot. 53: 323 (1915). Type: Ecuador: sandy fields, Riobamba, November, 1858, Spruce, 5788. Syntypes: BM, K.

Argentina, Bolivia (?), Ecuador, Peru.

Note: Robinson (2006: 187) was of the opinion that H. maritimum Kunth was also a synonym of H. diversifolium. Robinson (2006: 189) likened his concept of H. diversifolium to the Peruvian, Bolivian and Argentinian H. obtusifolium Cav.,

Heterosperma diversifolium Kunth var. tenuisectum Griseb., Abh. Knigl. Ges. Wiss. G ttingen 24(1): 198 (March-April 1879); Symb. Fl. Argent. : 198 (1879) = Heterosperma tenuisectum (Griseb.) Cabrera

Heterosperma involucratum (Phil.) Reiche, Anales Univ. Chile 112: 150 (1903) = Heterosperma nanum (Nutt.) Sherff

*Heterosperma maritimum Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 193 (1818) = ?Heterosperma ovatifolium Cav. see also comment under H. diversifolium!

Heterosperma nanum (Nutt.) Sherff, Bot. Gaz. 91: 312 (1931). [corrected to nanum]

Microdonta nana Nutt., Trans. Amer. Phil. Soc., ser. 2, 7: 370 (1841). Type: Hab. Near Arequipa, Peru. (Mr. Curson.) ... I have described from two specimens.

Bidens involucratum Phil., Anales Mus. Nac. Chile, Secc. 2, Bot. 8: 49 (1891). Type: [Chile: ] Calcalhuay in provincia Tarapac. Holotype: SGO.

Heterosperma pinnatum Cav. var. b involucratum (Phil.) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898).

Heterosperma involucratum (Phil.) Reiche, Anales Univ. Chile 112: 150 (1903).

Argentina, Bolivia (?), Chile, Peru.

Heterosperma ovatifolium Cav., Descr. Pl. : 204 (1802). [published as ovatifolia but corrected to ovatifolium]

Heterospemum ovatum Willd., Sp. Pl. 3: 2129 (1804).

*Heterosperma maritimum Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 193 (1818). Type: YT'Crescit in maritimis Peruvi, prope Truxillo. 0 Floret Septembri. Holotype: P-Bonpl.

Bidens (Heterospermum) rhombifolium Sch.Bip., Bonplandia 4(4): 54 (1856), nom. nud. (based on Lechler 1576).

*Heterospermum rhombifolium Griseb., Abh. K nigl. Ges. Wiss. Gttingen 19([1]): 186 (1874); Pl. Lorentz.: 138 (1874). Types: Tucuman, ad vias regionis subtropicae pr. Siambon. (Peruvia).

Heterospema pinnatum Cav. var. z ovatifolium (Cav.) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898).

Heterosperma pinnatum Cav. var. h maritimum (Kunth) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898).

Argentina, Bolivia (?), Ecuador, Peru.

Note: See comment under H. diversifolium vide H. maritimum

*Heterosperma pinnatum Cav., Icon. 3: 34 (1795). Type: .'Habitat in Nova-Hispania. 0 Florebat in Regio horto Matritense mense Octobri 1794. Note: There is much material in MA corresponding to this species seven sheets. MA (475793 Fiche 38/A6) with two almost identical handwritten labels Heterosperma pinnata/Cav. Ic. 2a 267/Heterospermum pinnatum Willd. Syng. pa. 2120/ex Hort. Reg. Matr./anno 1808, and one typewritten label Heterosperma pinnata Cav.; MA (475793 [sic!] a duplicate sheet Fiche 38/A7), with only a typewritten label as Heterosperma pinnata Cav.; MA (475798 Fiche 38/A8), with only a typewritten label as Heterospermum pinnatum Cav.; MA (475794 Fiche 38/B1) has a handwritten label Heterosperma pinnata/Icon./Queretaro in Nova-Hispan./Ne dedit. and one typewritten label ''Heterosperma pinnata Cav./Nueva Espaa; MA (475795 Fiche 38/B2) has a handwritten label '-'Heterosperma pinnata Cav./Ne Iter. and a typewritten label identifying the species; MA (475796 '- Fiche 38/B3) has two handwritten labels, the upper Heterosperma pinnata/Icon. t. 267/Sept. 1794 ex H. R. M. the lower Heterosperma/ex Queretaro in Nova-Hisp./Ne dedit. and one typewritten label (Heterosperma pinnata Cav./Nueva Espa a; MA (475797 Fiche 38/B4) has one handwritten label Heterosperma/Ixmiquiloan in Nova-hispania/Ne dedit.

Note: Foster (1958: 211) recorded this species, a native of Mexico, but it remains to be seen if the species does occur in Bolivia or if this refers to another.

*Heterosperma pinnatum Cav. [var.] g biternatum Kuntze, Revis. Gen. Pl. 3(3): 158 (1894) = H. tenuisectum (Griseb.) Cabrera

Heterosperma pinnatum Cav. var. d depressum (Griseb.) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898) = H. diversifolium Kunth

Heterosperma pinnatum Cav. var. e diversifolium (Kunth) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898) = H. diversifolium Kunth

Heterosperma pinnatum Cav. var. b involucratum (Phil.) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898) = Heterosperma nanum (Nutt.) Sherff

Heterosperma pinnatum Cav. var. h maritimum (Kunth) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898) = Heterosperma ovatifolium Cav.

Heterospema pinnatum Cav. var. z ovatifolium (Cav.) Kuntze, Revis. Gen. Pl. 3(3): 158 (1898) = Heterosperma ovatifolium Cav.

*Heterosperma rhombifolium Griseb., Abh. K nigl. Ges. Wiss. Gttingen 19([1]): 187 (1874) = H. ovatifolia Cav.

Heterosperma spathulatum S. F. Blake, J. Bot. 53: 323 (1915) = H. diversifolium (according to Robinson, 2006: 187).

Heterosperma tenuisectum (Griseb.) Cabrera, Fl. Prov. Jujuy 10: 404 (1978). [published as tenuisecta but corrected to tenuisectum]

Heterosperma diversifolium Kunth var. tenuisectum Griseb., Abh. K nigl. Ges. Wiss. Gttingen 24(1): 198 (March-April 1879); Symb. Fl. Argent. : 198 (1879). Types: [Argentina] Ct. Syntypes: Lorentz 562, Schickendantz 93, GOET.

Heterosperma pinnatum Cav. [var.] g biternatum Kuntze, Revis. Gen. Pl. 3(3): 158 (1898). Type: .'Bolivia: Tiraqui. [Bolivia. Tiraqui, 3600 m, 14 Apr 1892, Kuntze s.n. (2 sheets). according to Wetter & Zanoni, 1985: 333]

Argentina, Bolivia (Cochabamba).

Heterothalamus acaulis Wedd. ex R. E. Fr., Nova Acta Regiae Soc. Sci. Upsal., ser. 4, 1: 79 (1905) = Baccharis acaulis (Wedd. ex R. E. Fr.) Cabrera

Heterothalamus boliviensis Wedd., Chloris Andina 1: 179 (1856) = Baccharis boliviensis (Wedd.) Cabrera

Heterothalamus boliviensis Wedd. var. latifolius R. E. Fr., Nova Acta Regiae Soc. Sci. Upsal., ser. 4, 1: 78 (1905) = Baccharis boliviensis (Wedd.) Cabrera

Heterothalamus trinervis (Pers.) Hook, & Arn., J. Bot. (Hooker) 3(No. 17): 43 (Oct. 1840) = Baccharis trinervis Pers.

*Heterotheca deltoidea Klatt, Ann. Naturh. Hofmus. Wien 9: 359 (1894) = Munnozia hastifolia (Poepp.) H. Rob. & Brettell