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Gochnatia Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 15 (1818).

? Anastraphia D. Don, Trans. Linn. Soc. London 16(2): 295 (1830). Type: Anastraphia ilicifolia D. Don = Gochnatia ? Note: Although Cabrera (1971) discussed the problem of the application of Dons name he did not propose to which species of Gochnatia it should be referrable. The apparent absence of the Fraser material makes attribution of Dons genus to any modern concept somewhat difficult (vide Freire et al., 2002: 549).

Seris Less., Linnaea 5: 253 (1830), p.p. Type: not designated (but S. discoidea Less. now referrable to Richterago Kuntze).

Spadonia Less., Syn. Gen. Comp. : 99 (1832), p.p. Type: not designated, non Fries [FUNGI?]

Moquinia DC., Prodr. 7: 22 (1838), pp., non Spreng.f. (1828)(= Mouqiniella Balle [LORANTHACEAE])

Type: Gochnatia vernonioides Kunth

References

Beltran, H. & R. Ferreyra. (2001). Uma especie nueva de Asteraceae para Peru y Bolivia: Gochnatia lanceolata. Compositae Newslett. 36: 2630.

Cabrera, A. L. (1950). Observaciones sobre los gneros Gochnatia y Moquinia. Notas Mus. La Plata, Bot. 15(No. 74): 3748 & Figs. 1閖5.

Cabrera, A. L. (1971). Revisin del g nero Gochnatia (Compositae). Revista Mus. La Plata, Secc. Bot. 12(66): 1160.

Freire, S. E., Katinas, L. & G. Sancho. (2002). Gochnatia (Asteraceae, Mutisieae) and the Gochnatia complex: taxonomic implications from morphology. Ann. Missouri Bot. Gard. 89(4): 524550.

Sancho, G. (2000). Revisin y filogenia de la secci n Moquiniastrum Cabrera del gnero Gochnatia Kunth (Asteraceae, Mutisieae). Fontqueria 54(5): 61122.

Sancho, G., Freire, S. E., Katinas, L. & M. C. Teller a. (2005). A new combination and a new species of Andean Mutisieae (Asteraceae). TAXON 54(1): 8590.

Key to species

1. Inflorescence of solitary capitula 2

Inflorescence of many capitula 5

2. (1) Involucres cylindrical 3

Involucres campanulate; florets >12 4

3. (2) Florets 78; leaves 511 1.53.5 mm G. cardenasii

Florets c. 20; leaves 3545 613 mm G. angustifolia

4. (2) Involucrs 812 mm tall; florets 220, corollas 10 mm long G. curviflora

Involucres 1215 mm tall; florets >40, corollas 1215 mm long G. boliviana

5. (1) Plants monoecious G. palosanto

Plants dioecious 6

6. (5) Adult leaves pubescent on both surfaces G. rusbyana

Adult leaves pubescent beneath 7

7. (6) Pedicels ebracteolate; leaf venation not prominent G. polymorpha

Pedicels bracteolate towards involucre; leaf venation prominent beneath G. pulchra

Gochnatia angustifolia Sancho, S. E. Freire & Katinas, TAXON 54(1): 89 (2005). Type: Bolivia, Depto. Potos, Prov. Jos M. Linares Lizarazu, Jatum Palmar, 2750 m, 5 Apr 1993, Torrico & Peca 336. Holotype: LPB; isotype: US.

Bolivia (Potos).

*Gochnatia boliviana S. F. Blake, Contr. U.S. Natl. Herb. 22: 651 (1924). Type: ... collected at Santa Cruz, Bolivia, altitude 1,600 meters, May, 1892, by Otto Kuntze.󖑒 Holotype: US (701956); isotype: NY.

Bolivia (Cochabamba, Santa Cruz).

Note: Cabreras comments (1971: 25) suggested that other than size differences there is little difference between G. boliviana and G. curviflora and that the former may simply be just a multiflorous variety of the latter. There is apparently some geographical difference. Cabreras drawing suggests that the corolla tube is pubescent in this species and not in G. curviflora.

*Gochnatia cardenasii S. F. Blake, J. Wash. Acad. Sci. 25(7): 324 (1925). Type: Bolivia: Quechisla, Dept. Potos, alt. 3420 m., Dec. 1932, M. C rdenas 326. Holotype: US (1616163).

Argentina, Bolivia (Potos).

*Gochnatia curviflora (Griseb.) Hoffm. in Engl. & Prantl, Nat. Pflanzenfam. 4(5): 337 (1893).

Moquinia curviflora Griseb., Abh. K nigl. Gess. Wiss. Gttingen 24(1): 211 (March-April 1879); Symb. Fl. Argent. : 211 (1879). Type/s: O.: Tarija, Cuesta del Tambo. Cuesta de Buyuyu.

Argentina, Bolivia (Tarija).

Gochnatia genistoides (D. Don) Hook. & Arn., Companion Bot. Mag. 1(No. 4): 109 (1835) = Cyclolepis genistoides D. Don

Gochnatia lanceolata Beltran & Ferreyra, Compositae Newslett. 36: 26 (2001) = Chucoa lanceolata (Beltran & Ferreyra) Sancho, Freire, Katinas & Teller a (as to type), and Gochnatia angustifolia Sancho, S. E. Freire, Katinas (as to paratype)

*Gochnatia macrocephala (Rusby) Cabrera, Notas Mus. La Plata, Bot. 15(No. 74): 41 (1950) = Llerasia lucidula (S. F. Blake) Cuatrec.

Gochnatia malmei Cabrera, Notas Mus. La Plata, Bot. 1: 61 (1935) = Gochnatia polymorpha (Less.) Cabrera

Gochnatia palosanto Cabrera, Notas Mus. La Plata, Bot. 1(No. 3): 62 (1935). Type: [Argentina: ] Tucumn: Vipos, 786 m. s. m., Dep. Trancas, leg. C. Schreiter, no 85, 9-XI-1921. Holotype: C.

Argentina, Bolivia (Chuquisaca, Santa Cruz), Paraguay.

Palo santo (Cabrera, 1935: 63; Sancho, 1996: 11).

Gochnatia polymorpha (Less.) Cabrera, Notas Mus. La Plata, Bot. 15 (No. 74): 43 (1950).

Baccharis tomentosa Thunb., Pl. Bras. Dec. 3: 38 (1821), non Pers. (1807). Type: not cited. Cabrera (1950: 117) noted that the holotype was in UPS and that the material was s.loc., s.coll.

Spadonia polymorpha Less., Syn. Gen. Compos. : 101 (1832), taken as nom. nov. pro Baccharis tomentosa Thunb.

Spadonia polymorpha Less. [var.] a ceanothifolia Less., Syn. Gen. Compos. : 102 (1832). Type: not cited.

Spadonia polymorpha Less. [var.] b elaeagnifolia Less., Syn. Gen. Compos. : 102 (1832). Type: not cited.

Spadonia polymorpha Less. [var.] g obtusifolia Less., Syn. Gen. Compos. : 102 (1832). Type: not cited.

Spadonia polymorpha Less. [var.] d populifolia Less., Syn. Gen. Compos. : 102 (1832). Type: not cited.

Baccharis lessingiana DC., Prodr. 5: 414 (1836). Type: 椉L in Brasili. Bacch. nov. sp. Less.! mss. in Lund. pl. miss. .... (v. s. comm. cl. Lund.). Holotype: G-DC.

Moquinia polymorpha (Less.) DC., Prodr. 7: 23 (1838).

Moquinia polymorpha (Less.) DC. a ceanothifolia (Less.) DC., Prodr. 7: 23 (1838).

Moquinia polymorpha (Less.) DC. b elaeagnifolia (Less.) DC., Prodr. 7: 23 (1838).

Moquinia polymorpha (Less.) DC. g obtusifolia (Less.) DC., Prodr. 7: 23 (1838).

Moquinia polymorpha (Less.) DC. d populifolia (Less.) DC., Prodr. 7: 23 (1838).

Gochnatia malmei Cabrera, Notas Mus. La Plata, Bot. 1: 61 (1935). Type: Argentina. Misiones: Loreto, G. Grner 334, 21-I-1932. Holotype: LP; isotype: K.

Bolivia (Santa Cruz), Brazil, Paraguay.

Note: Lessings provision of the name Spadonia polymorpha is interpreted as having provided a new name for Thunbergs, since this was a later homonym (cf. Persoon, 1807 B. tomentosa (Ruiz & Pav.) Pers.). Lessing (1832: 101), however mentioned that there were numerous Sellow collections on which he based his description/s. None were linked directly to any of his varieties even though Cabrera (1950) and Sancho (2000) cite types for them. There is no indication that Cabrera saw type material during his travels to European herbaria in 1958. Both Cabrera (1950) and Sancho (2000) provided treatments dividing this species into a number of varieties, either of which can be used if an infraspecific level determination is needed.

Cambar (Cabrera & Klein, 1973); Cambara guasu, Kaa mar guasu (Cabrera, 1998).

Gochnatia pulchra Cabrera, Revista Mus. La Plata, Secc. Bot. 12(66): 106 (1971). Type: Brasil. S o Paulo: Ityrapina, leg. G. Gehrt [8296], 28-IV-1923. Holotype: LP; isotype: SP (8296), US.

Bolivia (Santa Cruz), Brazil.

*Gochnatia rusbyana Cabrera, Notas Mus. La Plata, Bot. 15(No. 74): 41 (1950), nom. nov. pro Moquinia boliviana Rusby

*Moquinia boliviana Rusby, Bull. New York Bot. Gard. 4(14): 399 (1907), non Gochnatia boliviana S. F. Blake. Type: [Bolivia:] ([Bang] No. 2252.) Holotype US (1403702); isotypes BM, CORD, G, K, LP, MO, M, US, W, Z (000003681).

Bolivia (Chuquisaca, La Paz, Santa Cruz), Peru.

Grangea domingensis (Cass.) M. Gmez [de la Maza] var. viscosa (L.) M. G mez [de la Maza], Dict. Bot. Nom. Vulg. Cubanos Puerto Riquenos : 115 (1889) = Egletes viscosa (L.) Less.