This page has been extracted from PDF and some formating may have been lost.
To see the original document click here


 

Culcitium Humb. & Bonpl., Pl. Aequinoct. 2: 1 (1808).

Senecio sect. Culcitium (Humb. & Bonpl.) Cuatrec., Fieldiana, Bot. 27(1): 50 (1950).

Type: not stated. Lectotype (selected by Cuatrecasas, 1951: 73): Senecio canescens (Humb. & Bonpl.) Cuatrec. = Culcitium canescens Humb. & Bonpl. Note: Culcitium rufescens Humb. & Bonpl. is cited as the type in Index Nominum Genericorum which suggested lectotypification, although the source is not indicated. Cabrera (1985: 112) simply cited C. rufescens as the type of the series under Senecio. Cuatrecasas (1951: 73) in transferring all species of Culcitium to Senecio Sect. Culcitium, noted Senecio canescens (Humb. & Bonpl.) Cuatrec. as the type of the section, with C. rufescens as a synonym of C. rufescens, which is reflected below; this is taken as the lectotypification of Culcitium.

Note: Rather than recognize Culcitium as a separate genus, merely as an altitudinal variant of more typical Senecio, Culcitium, Aetheolaena and Lasiocephalus have been considered by some authors best subsumed into Senecio sens. lat. (q.v. Pelser et al., 2007: 1098). This ignores the admitted usefullness of distribution, habit, corolla pigment, involucre, etc. in maintaining their recognition as separate genera. The inclusion of C. humile DC. is still doubtfull, especially with the rather variable leaves see comments below. Species such as C. humile might also key out to Senecio in the generic key to the Senecioneae, merely because of the less than woolly indumentum.

References

Cuatrecasas, J. (1950). The genus Culcitium H. et B. In: Studies on Andean Compositae I. Fieldiana, Bot. 27(1): 4051.

Cuatrecasas, J. (1951). Some Andean sections of the genus Senecio. In Studies on Andean Compositae II. Fieldiana, Bot. 27(2): 7074.

Note: Senecio comosus Sch.Bip. is excluded from Culcitium because it has radiate capitula. Senecio culcitioides Sch.Bip., later transferred by Cabrera (1985: 116) as S. comosus Sch.Bip. var. culcitioides (Sch.Bip.) Cabrera needs further investigation, especially to determine if it is conspecific with S. comosus, as suggested by Cabrera.

Culcitium albifolium Zoellner, Anales Mus. Hist. Nat. Valparaiso 3: 65 (1970). Type: Chile, Prov. de Tarapac, Depto. Arica en Portezuelo de Chapiqui a, a 4.500 m. de altura s. n. m.. Leg. Otto Zoellner Sch., 21 Enero 1969. Holotype: L.

Senecio zoellneri Martic. & Quezada, Bol. Soc. Biol. Concepcin 48: 102 (1974), nom. nov. pro Culcitium albiflorum Zoellneri

Bolivia (La Paz), Chile.

Culcitium boyacense Cuatrec., Caldasia 1: 5 (1940) = Culcitium canescens Humb. & Bonpl.

Culcitium canescens Humb. & Bonpl., Pl. Aeqin. 2: 4, t. 67 (1808/9). Type: Habitat in montosis Peruviae frigidis, frequens praesertim in monte Gualgayaoc. Holotype: P-Bonpl.

*Senecio canescens (Humb. & Bonpl.) Cuatrec., Fieldiana, Bot. 27(1): (1950).

Culcitium canescens Humb. & Bonpl. var. b monocephalum Wedd., Chloris Andina 1: 138 (1856). Types: '''C. nivale Schultz Bip., in Bonplandia, ann. 1856, p. 55, non H.B.K. Andes de Carabaya, prs de S. Antonio! h. 5000 m tr. (Lechler, exsicc., n. 1806); Cordillres de Guanaco (Haenke, dapr s DC.). Syntypes: P.

Culcitium boyacense Cuatrec., Caldasia 1(1): 5 (1940). Type: [Colombia:] Cordillera Oriental, Departamento de Boyac; Sierra Nevada del Cocuy, o de G icn, Quebrada de San Paulino, El Morr n, pramo 3800 m. alt., J. Cuatrecasas & H. Garc a Barriga, 11 sept. 1938 legerunt (no 1372). Holotype: COL; isotype: US (1797218).

Senecio boyacensis (Cuatrec.) Cuatrec., Fieldiana, Bot. 27(1): 43 (1950).

Senecio canescens (Humb. & Bonpl.) Cuatrec. var. boyacensis (Cuatrec.) Cuatrec., Fieldiana, Bot. 27(2): 36 (1951).

Senecio canescens (Humb. & Bonpl.) Cuatrec. var. macrocephalus Cuatrec., Fieldiana, Bot. 27(2): 36 (1951). Type: Colombia, Dept. Caldas, Nevado de Santa Isabel arenales 4420 m. alt., collect. J. Cuatrecasas 23208. Holotype: F.

Senecio canescens (Humb. & Bonpl.) Cuatrec. var. elongatus Cuatrec., Fieldiana, Bot. 27(2): 37 (1951). Type: Ecuador, Carchi: Pramo del Angel, 3300-3800 m. alt., collect. Acosta-Sol s 10562. Holotype: F.

Senecio canescens (Humb. & Bonpl.) Cuatrec. var. elongatus Cuatrec. f. magnificus Cuatrec., Fieldiana, Bot. 27(2): 37 (1951). Type: Ecuador, Carchi, Pramo del Angel 12,000 ft., collect. Edward Balls 7337. Holotype: US.

Senecio canescens (Humb. & Bonpl.) Cuatrec. var. ecuadoriensis Cuatrec., Fieldiana, Bot. 27(2): 37 (1951). Types: ''Ecuador, Monte Pichincha near Quito 4100-4500 m., collect. Hitchcock 21059 (US, NY).

*Senecio canescens (Humb. & Bonpl.) Cuatrec. var. monocephalus (Wedd.) Cuatrec., Fieldiana, Bot. 27(2): 37 (1951).

Bolivia (La Paz, Oruro), Colombia, Ecuador, Peru.

Note: Cuatrecasas (1951: 3637) suggested that there were few differences separating Culcitium canescens and C. rufescens Humb. & Bonpl. combining their synonymy and adding several new geographical varieties from Colombia and Ecuador. If C. rufescens is found to be the lectotype of Culcitium then appropriate changes will have to be made with the synonymy listed above.

Culcitium canescens Humb. & Bonpl. var. b monocephalum Wedd., Chloris Andina 1: 138 (1856) = Culcitium canescens Humb. & Bonpl.

Culcitium glaciale Meyen & Walp., Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19, Suppl. 1: 278 (1843) = Culcitium humile DC.

Culcitium humile DC., Prodr. 6: 325 (1838). Type: K inter Regiomontanas plantas herb. Haenkeani observ. ... (v.s. in h. Haenke cl. de Sternberg miss.) Holotype: PR; isotype: G-DC. Note: Cabrera (1985: 113) cited the holotype as in M, with an isotype in P!

Culcitium glaciale Meyen & Walp., Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19, Suppl. 1: 278 (1843). Type: 'Peruvia: in planitie circa Tacoram, alt. 1417,000 ped. (v.s.). Holotype: B.

*Senecio modestus Wedd., Chloris Andina 1: 105 (1856). Types: Hab. Prou: Cordill res de Carabaya!, prs de la limite inf rieure des neiges, sur les rochers humides (Wedd.). Bolivie: au sommet de la Cordillre de La Paz! (dOrbigny, no. 335).'' Syntypes: P. Lectotype (selected by Cabrera, 1985: 114): dOrbigny 355, P.

Senecio candollii Wedd., Chloris Andina 1: 106 (1856). Type: Hab. Perou? (Haenke). Holotype: ?P; isotype: ?PR.

Senecio mandonianus Wedd., Chloris Andina 1: 228 (1857). Type: Hab. Bolivie!: Cordillres du d partement de La Paz (Mandon). [Cited as Bolivia. Prov. Larecaja, Vicin. Sorata, leg. Mandon 109, 1857 by Cabrera (1985: 113)]. Holotype: P; isotypes: G, K, LP, S, US. Note: The Kew sheet of Mandon 109 has two specimens of Chersodoma antennaria (Wedd.) Cabrera mixed in with the Culcitium.

Senecio anacephalus Griseb., Abh. Knigl. Ges. Wiss. G ttingen 24(1): 208 (1879). Type: [Argentina] S.: Nevado desl Castillo, alt. 1015000'. [Lorentz & Hieronymus 91] Holotype: GOET; isotypes: CORD, LP.

Senecio glacialis (Mey. & Walp.) Cuatrec., Fieldiana, Bot. 27: 46 (1950).

Senecio modestus Wedd. var. candollii (Wedd.) H. Beltrn & Gal n de Mera, Bot. Complutensis 21: 104 (1996)

Argentina, Bolivia (Cochabamba, La Paz, Oruro, Potos), Peru.

Note: Usually treated as Senecio candollii Wedd. (e.g. Cabrera, 1985: 112), or with S. modestus Wedd. as a separate species, combined, under Culcitium, the earliest specific name is Culcitium humile DC. I very much doubt that this species should be kept in Culcitium since the leaves are at best pseudopetiolate with an elliptic lamina, and in some extreme cases, in Peruvian material (Tupayachi 6387), have very coarsely serrate, almost lobed, margins; the involucres have at best biseriate phyllaries with many calycular bracts. In the key to series of Senecio below (modified from Cabrera, 1985), material would most likely key to Ser. Crassicephali, although the indumentum is clearly different.

Culcitium lechleri Sch.Bip., Bonplandia 4(4): 55 (1856), nom. nud. = Chersodoma antennaria (Wedd.) Cabrera

*Culcitium neaei Sch.Bip. ex Wedd., Chloris Andina 1 :140 (1856). Type: Culcitium Neaei Schultz Bip., mscr. in herb. propr. YT- C. nivale, Neaei DC., Prodr., VI, 325. Hab. Prou: Cordill res de Guanoco (Haenke).

Culcitium nivale Kunth [var.] b? neaei DC., Prodr. 6: 325 (1838). Type: K in itin, per, Americ. legit cl. Ne. ... (v.s.) Holotype: G-DC.

Senecio neeanus Cuatrec., Fieldiana, Bot. 27(1): 44 (1950), as nom. nov. pro Culcitium neaei Sch.Bip. ex Wedd.

Bolivia (''Cerro Jurackasa, altovalle de Viloco), Peru.

*?Culcitium nivale Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. 4 (ed. folio): 134 (1818). Type: Crescit inter nives in monte ignivomo Antisan qua parte Chussulongum spectat, alt. 2340 hex. K Floret Martio. [Humboldt & Bonpland ''mss. n. 2249.] Holotype: P-Bonpl. Note: Foster (1958: 207) included this species for Bolivia, based on Cabreras record (1952: 197) for Herzog 2230, now considered to be Culcitium neaei, cf. Cabrera (1985: 115). Culcitium nivale is only known from Ecuador.

Culcitium nivale Kunth var. neaei DC., Prodr. 6: 325 (1838) = Culcitium neaei (DC.) Sch.Bip. ex Wedd.

Culcitium pflanzii Perkins, Bot. Jahrb. Syst. 49: 229 (1913) = Senecio pflanzii (Perkins) Cuatrec.

Culcitium serratifolium Meyen & Walp., Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19, Suppl. 1: 278 (1843) = Senecio serratifolius (Meyen & Walp.) Cuatrec.

Cyanopis Blume ex DC., Prodr. 5: 69 (1836), nom. illegit. superfl. non Cass. (1817) = Vernonia Schreb.