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Chromolaena DC., Prodr. 5: 133 (1836).

Eupatorium L. sect. Cylindrocephala DC., Prodr. 5: 141 (1836). Type: not stated. Lectotype: Eupatorium odoratum L. = Chromolaena odorata (L.) R. M. King & H. Rob.

Osmia Sch.Bip., Jahresber. Pollichia 22-24: 251 (1866). Type: Eupatorium odoratum L. = Chromolaena odorata (L.) R. M. King & H. Rob.

Heterolaena Sch.Bip. ex Benth. & Hook. f., Gen. Pl. 2: 245 (1873), nom. superfl. (based on Osmia Sch.Bip.)

Eupatorium L. sect. Osmia (Sch.Bip.) Benth. ex Baker in Mart., Fl. Bras. 6(2): 275 (1876).

Eupatorium L. sect. Chromolaena (DC.) Benth. ex Baker in Mart., Fl. Bras. 6(2): 300 (1876).

Eupatorium L. sect. Heterolaena (Sch.Bip. ex Benth. & Hook.f.) Baker in Mart., Fl. Bras. 6(2): 328 (1876).

Type: Chromolaena horminoides DC.

References

Gautier, L. (1992). Taxonomy and distribution of a tropical weed: Chromolaena odorata (L.) R. King & H. Robinson. Candollea 47: 645662.

King, R. M. & H. Robinson. (1970). Studies in the Eupatorieae (Compositae). XXIX. The genus Chromolaena. Phytologia 20(3): 196209.

King, R. M. & H. Robinson. (1982). Studies in the Eupatorieae (Asteraceae). CCXIV. New species of Chromolaena and Stevia from Bolivia. Phytologia 51(3): 172178.

*Eupatorium phyllocephalum Klatt, Ann. K. K. Naturhist. Hofmus. 9: 358 (1894). Types: Hab.: Bolivia (Chuquisaca), leg. dOrbigny, Nr. 1226, leg. Cuming. = Chromolaena sp.? Listed by Foster (1958: 209) but unplaced by King & Robinson (1987). Syntype: DOrbygny 1226 GH 2 (7898 & 7900).

Chromolaena alternifolia Gardner, London J. Bot. 5: 465 (1846) = Chromolaena stachyophylla (Spreng.) R. M. King & H. Rob.

Chromolaena arnottiana (Griseb.) R. M. King & H. Rob., Phytologia 20(3): 198 (1970).

*Eupatorium arnottianum Griseb., Abh. Knigl. Ges. Wiss. G ttingen 19([1]): 167 (1874); Pl. Lorentz.: 119 (1874). Types: Cordoba, infrequens in humidis pr. Las Peas, S. Francisco. (Tucuman, Entre Rios-Brasil. austr.)񻫒. Syntype: Lorentz, 178, GOET.

Argentina, Bolivia (Santa Cruz, Tarija).

Uou (Freire et al., 2006).

Chromolaena austera (B. L. Rob.) R. M. King & H. Rob., Phytologia 20(3): 199 (1970).

*Eupatorium ( Cylindrocephalum) austerum B. L. Rob., Contr. Gray Herb. 68: 9 (1923). Type: 駑Bolivia: in the bush region of Tres Cruces, alt. 1500 m., Feb. 1911 [Herzog 1610]. Holotype: G; isotypes: GH (photo and fragm.), L.

Bolivia (?).

Chromolaena bangii (Rusby) R. M. King & H. Rob., Phytologia 20(3): 199 (1970).

*Eupatorium bangii Rusby, Mem. Torrey Bot. Club 6(1): 56 (1896). Types: [Bolivia:] Vic. Cochabamba, 1891 ([Bang] 1133). Also collected in Bolivia by Bridges. Syntypes: K; isosyntype: Bang 1133, US. Note: Following Rusbys comments (Rusby, 1993) about the distribution of material and the assistance that Britton provided over notes on material not examined by Rusby, it is more than likely that the Bridges colleciton was seen only at K; the slip of paper pinned to the Bridges sheet in K supports this.

Bolivia (Cochabamba), Peru.

Chromolaena beckii R. M. King & H. Rob. Phytologia, 51(3): 172 (1982). Type: BOLIVIA: Santa Cruz: Prov. A. Ibaez, Santa Cruz ca. 22 kms. hacia Abap (Sud). ca. 500 m.s.n.m. Chaparral abierto con cesped bajo. -20 cm. 14.3.1981. St. G. Beck 6502. Holotype: US; isotype: LPB

Bolivia (Santa Cruz).

Chromolaena connivens (Rusby) R. M. King & H. Rob., Phytologia 20(3): 200 (1970).

Eupatorium paucidentatum Sch.Bip., Bull. Soc. Bot. France 12: 81 (1865); Linnaea 34(5): 535 (Feb. 1866), nom. nud. (based on Mandon 248), non Sch.Bip. ex Baker (1876) (= Chromolaena congesta (Hook. & Arn.) R. M. King & H. Rob.)

*Eupatorium connivens Rusby, Mem. Torrey Bot. Club 6(1): 57 (1896). Types: [Bolivia:] Vic. Cochabamba, 1891 ([Bang] 1114). The same as Mandons 248. Also collected by Bridges in Bolivia. [Isosyntype (Bang 1114): US, Z (000003348).]

*Eupatorium cochabambense Hieron., Bot. Jahrb. Syst. 22(45): 745 (1897). Types: Bolivien: bei Cochabamba (O. Kuntze, 26. Mrz 1892); Mig. Bang, 1891, n. 1114 unter dem Namen E. paucidentatum Schultz-Bip. ausgegeben). Syntypes: Bퟅ.

Argentina, Bolivia (Cochabamba, La Paz, Santa Cruz), Peru.

Chromolaena desmocephala (B. L. Rob.) R. M. King & H. Rob., Phytologia 49(1): 4 (1981).

*Eupatorium ( Cylindrocephalum) desmocephalum B. L. Rob., Contr. Gray Herb. 68: 14 (1923). Type: Bolivia: on rocky slopes of the Cuesta de Guayabillas, alt. 1200 m., March, 1911, Th. Herzog, no. 1739. Holotype: G; isotype: GH (fragment).

Bolivia (Tarija).

Chromolaena extensa (Gardner) R. M. King & H. Rob., Phytologia 20(3): 201 (1970).

*Eupatorium extensum Gardner, London J. Bot. 6: 440 (1847). Type: [Brazil] Hab. Bushy places between Capella da Posse and San Pedro, Province of Goyaz. May, 1840. [Gardner] 4214.

Bolivia (La Paz, Santa Cruz), Brazil, Peru.

Britton (1891) cited Guanai, 2,000 ft. ([Rusby] 1627).

Chromolaena herzogii (B. L. Rob.) R. M. King & H. Rob., Phytologia 20(3): 201 (1970).

*Eupatorium ( Cylindrocephalum) herzogii B. L. Rob., Contr. Gray Herb. 68: 19 (1923). Type: Bolivia: common on mountain meadows near Samaipata, Dec. 1907, Th. Herzog, no. 694. Holotype: Z (000003355); isotype: GH (fragment).

Bolivia (Santa Cruz).

?Chromolaena hirsuta (Hook. & Arn.) R. M. King & H. Rob., Phytologia 20(3): 201 (1970).

*Eupatorium hirsutum Hook. & Arn., Companion Bot. Mag. 1(No. 8): 239 (1836). Type: Banda Orientale and Rio Grande do Sul, Tweedie. Note: There is no distinction between the typical variety and the unnamed variety b as it appears only one collection may have been cited.

Note: Although listed by Foster (1958: 208) under Eupatorium, King & Robinson (1987: 386) only listed the species from Argentina, Brazil and Uruguay.

Chromolena hookeriana (Griseb.) R. M. King & H. Rob., Phytologia 47(3): 231 (1980).

Eupatorium ciliatum Hook. & Arn., Companion Bot. Mag. 1(No. 8): 240 (1836), nom. illegit., non Less. (1831) (= Ageratina ciliata (Less.) R. M. King & H. Rob.). Type: Buenos Ayres, Tweedie. Holotype: K.

Eupatorium hookerianum Griseb., Abh. Knigl. Ges. Wiss. G ttingen 19([1]): 166 (1874), as nom. nov. pro E. ciliatum Hook. & Arn.

*Eupatorium conyzoides Vahl var. ciliatum (Hook. & Arn.) Hieron., Bot. Jahrb. Syst. 22(45): 741 (1897).

Eupatorium jujuiense Hieron., Bot. Jahrb. Syst. 22(45): 744 (1897). Type: Jujui: bei ser Stadt Jujui (Lor. u. Hieron., April 1873, n. 1017). Holotype: B.

Eupatorium hookerianum Griseb. var. jujuiense (Hieron.) Cabrera & Vittet, Revista Mus. La Plata, Secc. Bot. 8: 216 (1954).

Chromolaena jujuiensis (Hieron.) R. M. King & H. Rob., Phytologia 20(3): 202 (1970).

Argentina, Bolivia (La Paz).

Chromolaena ivifolia (L.) R. M. King & H. Rob., Phytologia 20(3): 202 (1970).

*Eupatorium ivifolium L., Syst. Nat. (ed. 10) : 1205 (1759). Type: Habitat in Jamaica. Lectotype (selected by King & Robinson in Woodson & Schery, 1975: 924): Browne, Herb. Linn. No. 978.28 (LINN).

Eupatorium concinnum Hook. & Arn., Companion Bot. Mag. 1(No. 8): 239 (1835), non DC. (1836) (= Neocabreria concinna R. M. King & H. Rob.). Types: Banda Orientale and Rio Grande, Tweedie. Syntypes: K.

Eupatorium obscurum DC., Prodr. 5: 142 (1836). Type: L in Brasiliae prov. Rio-Grande. ... (v. s. in h. Mus. reg. ar. Mus. imp. Bras. sine num. miss.). Holotype: P; isotype: G-DC.

Ooclinium clavatum Benth., Ann. Nat. Hist. 2(8): 108 (Oct. 1838). Type: ퟙBritish Guiana. Schomburgk, n. 165. Holotype: K. Note: The IPNI data base, Baker in Flora Brasiliensis and other references misquote the place of publication as Ann. Nat. Hist. 1(2): 108 (1839) apart from the wrong date, this is an interesting article by Mr Gunn on mammals and fishes of Van Diemens Land.

Osmia ivifolia (L.) Sch.Bip., Jahresber. Pollichia 22-24: 252 (1866).

Osmia obscura (DC.) Sch.Bip., Jahresber. Pollichia 22-24: 252 (1866).

Eupatorium ivifolium L. var. serratum Griseb., Cat. Pl. Cub.: 146 (1866). Type: [Cuba:] Wr[ight]. 2800. Holotype: GOET; isotype: K.

Eupatorium polyanthum Sch.Bip. ex Baker in Mart. Fl. Bras. 6(2): 285 (1876). Types: Habitat in Brasilia meridionali, loco acuratius non designato: Sello n. 511, 545; Uruguay ad ripas fluminis Rio Negro: Gibert n. 226. Syntype: Gibert 226, K 2.

Osmia polyantha Sch.Bip. ex Baker in Mart., Fl. Bras. 6(2): 285 (1876), nom. nud. pro syn.

*Eupatorium ivifolium L. var. b extrorsa [sic!] Baker in Mart., Fl. Bras. 6(2): 290 (1876). Types: [Brazil:] Ad Sorocaba prov. S. Paulo: Riedel; prov. Minas Geras ad Lagoa Santa: Warming. Syntype: Riedel s.n., K.

Eupatorium extrorsum Sch.Bip. ex Baker in Mart., Fl. Bras. 6(2): 290 (1876), nom. nud. pro syn.

Eupatorium ivifolium L. var. g gracillima Baker in Mart., Fl. Bras. 6(2): 290 (1876), based on Ooclinium clavatum Benth.

Osmia gracillima Sch.Bip. ex Baker in Mart., Fl. Bras. 6(2): 290 (187), nom. nud. pro syn.

Osmia stenophylla Sch.Bip. ex Baker in Mart., Fl. Bras. 6(2): 290 (1876), nom. nud. pro syn.

Eupatorium ivifolium L. var. d laxiflora Baker in Mart., Fl. Bras. 6(2): 290 (1876). Type: [Brazil:] 뒑Ad ripas fluminis Maranho: Pohl n. 289. Note: There are two sheets of this collection in K, the material ex herb. Benthamianum bearing the locality handwritten by Baker on the sheet; this sheet is taken as the holotype.

Eupatorium ivifolium L. var. e aspera Baker in Mart., Fl. Bras. 6(2): 291 (1876). Type: [Brazil:] 㒑Prov. Goyaz in udis ad Arrayas: Gardner n. 4207. Holotype: K (ex herb. Hookerianum).

*Eupatorium guanaiense Britton, Bull. Torrey Bot. Club 18(11): 333 (1891). Type: [Bolivia:] Guaniai, 2,000 ft. ([Rusby] 1735). Holotype: NY; isotype: US.

Eupatorium ivifolium L. var. genuinum Hassl., Feddes Repert. Spec. Nov. Regni Veg. 14(1015): 280 (1916).

Eupatorium ivifolium L. var. hirsutum Hassl., Feddes Repert. Spec. Nov. Regni Veg. 14(1015): 280 (1916). Types: Paraguay: Hassler no. 1518a, 6174. Pl. Hassl., l.c., Caaguaz, Hassler no. 9205 bis Estrella, Hassler no. 10267. Gran Chaco: Santa Elisa, Hassler no. 2602, 2788. Syntypes: G; isosyntypes: Hassler 2788 & 10267, K.

Eupatorium ivifolium L. var. pilcomayense Hassl., Feddes Repert. Spec. Nov. Regni Veg. 14(1015): 280 (1916). Type: [Paraguay:] Gran Chaco: Ad flumen Pilcomayo, Rojas no. 236.摒 Holotype: G.

Eupatorium ivifolium L. var. perturbinatum B. L. Rob., Contr. Gray Herb. 80: 23 (1928). Type: Argentina: Formosa: Guayalec, Jrgensen, no. 3351. Holotype: GH.

Osmia gracillima Sch.Bip. ex Malme, Ark. Bot. 24A(8): 23 (1932), nom. nud. pro syn.

SE United States, Greater Antilles, Lesser Antilles, Mexico, Central America, South America.

Bolivia (La Paz, Santa Cruz), Brazil.

Note: Eupatorium affine Hook. & Arn. was equated with Chromolaena odorata by King & Robinson (1987), although syntype material in K most certainly doesnt resemble that species. Baker (1876) clearly equated it with Ch. ivifolia.

Typycha pito (Cabrera, 1996); Eupatrio (Cabrera & Klein, 1991); Piron k-lek (Freire et al., 2006).

Chromolaena jujuiensis (Hieron.) R. M. King & H. Rob., Phytologia 20(3): 202 (1970) = Chromolaena hookeriana (Griseb.) R. M. King & H. Rob.

Chromolaena laevigata (Lam.) R. M. King & H. Rob., Phytologia 20(3): 202 (1970).

*Eupatorium laevigatum Lam., Encycl. 2: 408 (1786). Type: Cette plante nous a t communique par M. de Jussieu; nous la croyons originair dAm rique. (v.s.). Holotype (cited by King & Robinson, 1975: 924): P-JU (8394); isotype: P-LA (306/20). Note: Although King & Robinson cited the holotype as a Jussieu collection the original protologue citation indicates that Jussieu sent the material Lamarck suggesting that the material in P-LA is most likely to be considered the holotype, that in P-JU the isotype.

Eupatorium australe Thunb., Pl. Bras., Decas 2: 26 (1818). Type: not cited.

Chrysocoma punctata Vell., Fl. Flum. 8: 335, tab. 49 (1825)[7 Sept. - 28 Nov. 1829]. Type: Habitat fruticetis maritimis.

Eupatorium psiadiifolium DC., Prodr. 5: 144 (1836). Type: L in Brasili (h. LH r.!), in siccis ad Cercovado (Lhostky!). (v.s.). Syntype: Lhostky s.n., G-DC. Presumably the LHritier syntype is in NEU; there is collection in G-DC without a collectors name on the label which may represent a duplicate of this material.

Eupatorium psiadiifolium DC. var. b latifolium DC., Prodr. 5: 145 (1836). Types: ꨉL in Cajenn legit cl. Patris! (v. s.). Holotype: G-DC.

Eupatorium propinquum DC., Prodr. 5: 146 (1836). Type: ⒑L in Brasiliae prov. Rio-Grande. ... (v. s. in h. Mus. reg. Par. Mus. imp. Bras. miss. sub n. 1064.). Holotype: P; isotype: G-DC.

Eupatorium fasciculare Poepp., Nov. Gen. Sp. Pl. 3: 54 (1845). Type: ґCrescit in Peruviae subandinae montibus calcareis herbidis ad Cassapi. Januario lectum. Holotype: ?

Eupatorium resinosum Poepp., Nov. Gen. Sp. 3: 54 (1845). Type: Crescit cum praecedente. Floret Octobre. q.v. Eupatorium fasciculare Crescit in Peruviae subandinae montibus calcareis herbidis ad Cassapi. Januario lectum. Holotype: W.

Osmia alternans (DC.) Sch.Bip., Jahresber. Pollichia 22-24: 252 (1866).

Osmia laevigata (Lam.) Sch.Bip., Jahresber. Pollichia 22-24: 252 (1866).

Osmia propinqua (DC.) Sch.Bip., Jahresber. Pollichia 22-24: 252 (1866).

Eupatorium laevigata Lam. var. squamulosa Hieron., Bot. Jahrb. Syst. 22(4-5): 747 (1897). Type: Brazil: Santa Catharina: in der Capoeira bei So Francisco (Ule, M rz 1884, n. 136). Holotype: B; isotype: S.

Eupatorium laevigata Lam. var. submembranaceum Hieron., Bot. Jahrb. Syst. 22(4-5): 748 (1897). Type: [Argentina:] Salta: bei der Estancia Yacone (Lor.[entz] u. Hieron.[ymus], Mrz 1873, n. 325). Isotype: GOET.

*Eupatorium laevigatum Lam. f. albiflorum Kuntze, Revis. Gen. Pl. 3(3): 147 (1898). Type: 䒑Bolivia: Yapacani, 400 m. [Bolivia. Yapacani, 400 m, Jun 1892, Kuntze s.n. according to Wetter & Zanoni, 1985: 330]

*Eupatorium laevigatum Lam. f. flavidum Kuntze, Revis. Gen. Pl. 3(3): 147 (1898). Type: Bolivia: 1600 m Santa Cruz. [Bolivia. Sierra de Santa Cruz, 1600 m, May 1892, Kuntze s.n. according to Wetter & Zanoni, 1985: 331]

*Eupatorium laevigatum Lam. f. lilacinum Kuntze, Revis. Gen. Pl. 3(3): 147 (1898). Types: Bolivia: Ost-Velasco 200 m, Yapacani 400 m. [Bolivia. Ost-Velasco, 200 m, Jul 1892, Kuntze s.n. according to Wetter & Zanoni, 1985: 331].

Eupatorium laevigatum Lam. var. psidiifolia (DC.) Hassl., Feddes Repert. Spec. Nov. Regni Veg. 14(1015): 282 (1916).

Eupatorium laevigatum Lam. var. longepetiolatum Hassl., Feddes Repert. Spec. Nov. Regni Veg. 14(10-15): 282 (1916). Types: Gran Chaco: Ad ripas fluminis Paraguay, Hassler no. 2327, (foliis ovatis cuspidulato-acuminatis argute serratis): eod, loco no. 2327a. (foliis ovato-ellipticis apice obtusis breviter cuspidatis crenato-serratis). Syntypes: G.

Argentina, Bolivia (La Paz, Santa Cruz), Brazil, Mexico.

Sanatodo (Cabrera & Freire, 1997: 33), Cambar, Cambar -falso, Cambarazinho, Mata-pasto, Vassoura-do-campo (Cabrera & Klein, 1991); Ca-h , Campanela, Doctorcito (Freire et al., 2006).

Chromolaena leptocephala (DC.) R. M. King & H. Rob., Phytologia 20(3): 202 (1970).

*Eupatorium leptocephalum DC., Prodr. 5: 148 (1836). Type: L in Andibus Peruvianorum legit indef. Haenke. ... (v.s. in h. Haenke cl. Sternb. comm.). Holotype: PR; isotype: G-DC.

*Eupatorium leptocephalum DC. var. hypomalacum B. L. Rob., Contr. Gray Herb. 80: 24 (1928). Type: ґPeru: Urubamba Valley at Cedrobamba, 2200 m., Aug. 1927, Prof. F. L. Herrera, no. 1572. Holotype: GH. Note: the following paratype was also cited: Bolivia: Prov. Larecaja: in woods, Queliguaya, vicinity of Sorata, in temperate region, alt. 2700 m., Sept. 1858, Mandon, no. 247 G, GH, K, P.

Bolivia (La Paz), Colombia, Ecuador, Peru. The record for Bolivia, recorded by B.L. Robinson (1920) and based on a Mandon collection (Mandon 247) was originally determined by Schultz Bipontinus, and much later by Robinson himself, for the variety (q.v. Robinson, 1828: 24).

Chromolaena mallota (B. L. Rob.) R. M. King & H. Rob., Phytologia 20(3): 203 (1970).

*Euaptorium mallotum B. L. Rob., Proc. Amer. Acad. Arts 55: 22 (1919). Type: Bolivia: Dept. LaPaz [sic!], Prov. Larecaja: viciniis Sorata; inter Munaypata et rivum Chalassayo, in schistosis. Reg. temp. 26002700 m., Mandon, no. 249. Holotype: GH; isotypes: NY, S.

*Eupatorium mallotum B. L. Rob. var. aporum B. L. Rob., Proc. Amer. Acad. Arts 55: 23 (1919). Type: Bolivia without locality: Bang, no. 2875 (Gr., U.S.). Syntypes: GH, US.

Bolivia (La Paz).

Chromolaena odorata (L.) R. M. King & H. Rob., Phytologia 20(3): 204 (1970).

Eupatorium odoratum L., Syst., ed. 10, 2 : 1205 (1759). Type: Habitat in America. Lectotype (selected by King & Robinson in Woodson & Schery 1975: 925): [icon] Eupatoria Conyzoides folio molli & incano, capitulis magnis, Americana in Plukenet, Phytographia: t. 177, f. 3 (1692); Almag. Bot. : 141 (1696). Typotype: Herb. Sloane 96: 25 (BM-SL).

Eupatorium conyzoides Vahl, Symb. Bot. 3: 96 (1794), nom. illegit., non Mill. (1768) (= Vernonia arborescens (L.) Sw.). Type: Habitat in America meridionali? Dn. Thouin. Holotype: ?C.

Eupatorium floribundum Kunth in Humb., Bonpl. & Kunth., Nov. Gen. Sp. Pl. 4 (ed. folio): 92 (1818). Type: Crescit locis frondosis, temperatis Loxam inter et fluvium Catamayo, alt. 1080 hex. (Peruvia.) L Floret Augusto. [Humboldt & Bonpland mss. n. 3426. Gonzanama]. Holotype: P-Bonpl.

? Chrysocoma maculata Vell., Fl. Flum. : 325 (1825)[7 Sept. - 28 Nov. 1829]; Fl. Flum. Icones 8: tab. 6 (1831). Type: Habitat fruticetis maritimis Pharmacopolitanis. Floret April. May.

Eupatorium divergens Less., Linnaea 5: 138 (1830). Type: Prope Hacienda de la Laguna. Octbr. Holotype: B.

Eupatorium clematitis DC., Prodr. 5: 144 (1836). Type: L in Peruvi? legit cl. Poeppig. (v. s. comm. cl. inv. sub n. 3108.). Holotype: G-DC.

Eupatorium graciliflorum DC., Prodr. 5: 145 (1836). Types: L in Mexico leg. cl. Haenke, circa Acapulco (Karvinski). ... (v. s. in h. Haenke cl. Sternb. comm. et h. Acad. reg. Monac.). Syntypes: G-DC; isosyntype: Haenke s.n., M, PR.

Eupatorium conyzoides Vahl var. heterolepis Griseb., Fl. Brit. W.I. : 358 (1861) Types: ґHab. Bahamas!, Swains.; Jamaica! all coll. (a, b); Antigua!, Nichols., Wullschl.; [Cuba! and Mexico! to Brazil!]. Location of all syntypes unknown. Syntypes: Wullschlaegel 883, Wright, 294, GOET.

Osmia odorata (L.) Sch.Bip., Jahresb. Pollichia 22/24: 250 (1866).

Osmia divergens (Less.) Sch.Bip., Jahresb. Pollichia 22/24: 252 (1866).

Osmia graciliflorum (DC.) Sch.Bip., Jahresb. Pollichia 22/24: 252 (1866).

?Eupatorium conyzoides Vahl var. d incanum Baker in Mart., Fl. Bras. 6(2): 278 (1876). Types: [Brazil:] In prov. Bahia: Salzmann, Blanchet, Sello n. 565; prov. Mato Grosso ad Cuiab: Manso n. 91. Note: There are two collections of Salzmann 14 in K but neither were determined by Baker as var. incanum; they may well be snytype material. None of the other syntypes are apparently in K; the Manso collection is in BR.

Eupatorium conyzoides Vahl var. e pauciflorum Baker in Mart., Fl. Bras. 6(2): 278 (1876). Types: ᒑIn Brasilia meridionali: Riedel n. 358; Surinamia: Wallschlaegel n. 883. Frequens etiam in ditione occidentali Andina et in Antillia. Note: None of these collections, nor material annotated by Baker, has been found in K.

Eupatorium klattii Millsp., Publ. Field Columbian Mus., Bot. Ser. 2: 105 (1900). Type: Cuba, shores of Santiago Bay. Millspaugh 1126. Holotype: F.

Eupatorium odoratum L. var. pauciflorum (Baker) Hieron., Bot. Jahrb. Syst. 28(5): 564 (1901).

Eupatorium conyzoides Vahl var. scaberulum Hassl., Repert. Spec. Nov. Regni Veg. 14: 279 (1916). Type: Paraguay: Ad margines silvarum, scandens, Cordillera de Altos, Hassler no. 11859. Holotype: G.

SE USA, Mexico, West Indies south to Argentina and widely adventive. Bolivia (La Paz, Santa Cruz).

La Paz: Franz Tamayo, 688 m, 28 Septiembre 2005, Torrez et al. 3 (K, MO).

Kaa kat (Cabrera, 1996); Cruzeiro, Eupat rio (Cabrera & Klein, 1991); Pilarcito (Freire et al., 2006).

Chromolaena oxylepis (DC.) R. M. King & H. Rob., Phytologia 20(): 204 (1970).

Eupatorium oxylepis DC., Prodr. 5: 145 (1836). Type: L in Brasiliae prov. Sancti-Pauli. ... (v. s. in h. Mus. reg. Par. Mus. imp. Bras. miss. sub. n. 468.) Holotype: P; isotype: G-DC (fragments of leaves anda a short piece of stem).

Bolivia (Santa Cruz), Brazil.

Chromolaena porophylloides (B. L. Rob.) R. M. King & H. Rob., Phytologia 49(1): 4 (1981).

*Eupatorium (ҧ Praxelis) porophylloides B. L. Rob., Contr. Gray Herb. 68: 29 (1923). Type: Bolivia: Dept. of Santa Cruz: Prov. of Chiquitos: on plain of the Cerro de Santiago, alt. 800 m., May, 1907, Th. Herzog, no. 24. Holotype: Z (000003365); isotype: GH (fragment).

Bolivia (Santa Cruz).

Chromolaena pratensis Gardner, London J. Bot. 1: 176 (1842) = Campuloclinium macrocephalum (Less.) DC.

Chromolaena squalida (DC.) R. M. King & H. Rob., Phytologia 20(3): 206 (1970).

Eupatorium squalidum DC., Prodr. 5: 142 (1836). Type: L in Brasiliae, prov. Minarum Gener. ad Mariannum legit cl. Vauthier. ... (v. s. comm. cl. Vauthier sub n. 279.). Holotype: G-DC.

Eupatorium ramosissimum Gardner, London J. Bot. 6: 441 (1847). Type: [Brazil: ] Hab. Margins of woods near Villa de Arrayas, Province of Goyaz. March, 1840.ђ [Gardner] 4206.

Eupatorium crenatum Gardner, London J. Bot. 6: 441 (1846). Type: [Brazil: ] Hab. Near Villa de Arrayas, Province of Goyaz. April 1840. [Gardner] 3829. Holotype BM; isotypes K 2, .....

Eupatorium venosum Mart. ex Baker in Mart., Fl. Bras. 6(2): 281 (1876), nom. nud. pro syn.

Eupatorium sitiense Hieron., Bot. Jahrb. Syst. 22(4-5): 751 (1897). Types: [Brazil: ] Minas Geras: im Camp bei Sitio (H. Schenk 20. M rz 1887, n. 3184 und 3197). Syntypes: B. (S).

*Eupatorium squalidum DC. var. rusbyanum B. L. Rob., Proc. Amer. Acad. Arts 55: 34 (1919). Type: Bolivia: at Guanai in Prov. Laraceja, Dept. La Paz, alt. 610 m., May, 1886, Rusby, no. 1623. Holotype: NY; isotype: US.

Bolivia (Santa Cruz), Brazil, Guyana, Paraguay, Peru, Venezuela.

Chromolaena squarrosoramosa (Hieron.) R. M. King & H. Rob., Phytologia 20(3): 206 (1970).

*Eupatorium squarrosoramosum Hieron., Bot. Jahrb. Syst. 22(45): 753 (1897). Types: [Argentina:] Tucuman: im Camp bei Estancia La Cruz (Lor., 2024 April 1872, n. 187). Salta: in der Quebrada de San lorenzo unweit Salta (Lor. u. Hieron., 9. Mrz 1873). Syntypes: B䒆.

Hieronymus (1897) placed in synonymy E. squarrulosum Griseb., Plant. Lorentz. p. 119. n. 397, Symb. p. 168. n. 1003. partim, non Hook. & Arn..

Argentina, Bolivia (Tarija), Brazil.

Chromolaena stachyophylla (Spreng.) R. M. King & H. Rob., Phytologia 20(3): 206 (1970).

*Eupatorium stachyophyllum Spreng., Syst. Veg., ed. 16, 3: 420 (1826). Type: Brasil. Sello. Holotype: P.

Eupatorium subalternifolium DC., Prodr. 5: 152 (1836). Types: K in Brasiliae prov. Minarum General. ad Serra do Frio (Vauth.! n. 335), in campis editis prov. Sancti-Paulo (Lund herb. n. 851!). ... (v. s.) Syntypes: G-DC.

Eupatorium scaberrimum Walp., Linnaea 14: 505 (1840). Type: In Brasiliae Campos boa Perna legit Luschnath. Holotype: B.

Chromolaena alternifolia Gardner, London J. Bot. 5: 465 (1846). Type: [Brazil:] Hab. Dry hills near Morro Velho, Province of Minas Gerais, Brazil. Fl. in Sept. [Gardner] 4860.

Eupatorium pulchrum Gardner, London J. Bot. 6: 444 (1847), nom. superfl., pro Chromolaena alternifolia Gardner

Eupatorium claussenii Gardner, London J. Bot. 6: 445 (1847). Type: Hab. Province of Minas Geraes, Brazil. Claussen. Holotype: K.

Bolivia (?), Brazil.

Chromolaena subscandens (Hieron.) R. M. King & H. Rob., Phytologia 20(3): 207 (1970).

Eupatorium subscandens Hieron., Bot. Jahrb. Syst. 22(45): 742 (1897). Type: Bolivien: in der Nhe von Cochabamba (Mig. Bang, 1891, n. 1208, als E. conyzoides verteilt). Holotype: B䒆; isotypes:

Bolivia (Cochabamba).

Chromolaena toldensis (Hieron.) R. M. King & H. Rob., Phytologia 20(3): 207 (1970).

*Eupatorium toldense Hieron., Bot. Jahrb. Syst. 40(3): 378 (1908). Type: Bolivia: prope Toldos haud procul a pago Bermejo, alt. s. m. 1800 m, locis campestribus sparse herbis obsitis (K. Fiebrig n. 2371; 8. m. Dec. 1903). Holotype: B; isotype: S.

Argentina, Bolivia (Tarija).

Chromolaena tunariensis (Hieron.) R. M. King & H. Rob., Phytologia 20(3): 207 (1970).

E. conyzoides Vahl var. tunariensis Hieron., Bot. Jahrb. Syst. 22(45): 742 (1897). Types: Bolivien: bei Tunari um 2400 m ber Meer (O. Kuntze, April, Mai 1892); in der Sierra bei Santa Cruz (O. Kuntze, Mai 1892). Syntypes: B.

*Eupatorium tunariense (Hieron.) B. L. Rob., Contr. Gray Herb. 61: 39 (1920).

Argentina, Bolivia (Cochabamba, La Paz, Santa Cruz).