This page has been extracted from PDF and some formating may have been lost.
To see the original document click here
Chaptalia
Vent., Descr. Pl. Jard. Cels : tab. 61 (1802).
Thyrsanthema
Necker, Elem. Bot. 1: 6 (1790), nom. rej. Type: not designated. (See discussion
by Neson (1995: 154155)).
Leria DC., Ann.
Mus. Natl. Hist. Nat. 19: 68 (1812). Type: not stated. Lectotype (selected by
Burkart, 1944: 561). Type: Tussilago
nutans L. = Chaptalia nutans
(L.) Polak.
Loxodon Cass.,
Dict. Sci. Nat. 27: 253 (1823). Type: not state. Lectotype (selected by
Burkart, 1944: 534). Type: Tussilago
exscapa Pers. = Loxodon exscapa
(Pers.) Cass. = Chaptalia exscapa (Pers.) Baker
Lieberkuhna Cass.,
Dict. Sci. Nat. 27: 286 (1823). Lectotype (selected by Burkart, 1944: 539): Perdicium piloselloides Vahl = Chaptalia
piloselloides (Vahl) Baker
Oxydon Less.,
Linnaea 5: 357 (1830). Type: Oxydon
bicolor Less. = Chaptalia runcinata Kunth
Oxyodon DC., Prodr.
7: 43 (1838), orth. var. pro Oxydon
Less.
Chaptalia Vent.
sect. Archichaptalia Burkart,
Darwiniana 6(4): 525 (1944). Type: Chaptalia
rotundifolia D. Don
Chaptalia Vent.
sect. Pseudotrichocline Burkart,
Darwiniana 6(4): 524 & 532 (1944). Type: Chaptalia isernia Cuatrec.
Chaptalia Vent.
sect. Loxodon (Cass.) Burkart, Darwiniana
6(4): 524 & 534 (1944).
Chaptalia Vent.
sect. Lieberkuhna (Cass.) Burkart,
Darwiniana 6(4): 524 & 539 (1944).
Chaptalia Vent.
sect. Leria (DC.) Burkart, Darwiniana
6(4): 525 & 560 (1944). Lectotype (selected by Burkart, 1944: 561): Chaptalia nutans (L.) Polak.
Chaptalia Vent.
sect. Microchaptalia Burkart,
Darwiniana 6(4): 525 & 586 (1944), p.p. Type: Tussilago pumila Sw. = Chaptalia pumila (Sw.) Urban
Type: Chaptalia tomentosa Vent.
References
Burkart, A. (1944). Estudio del gnero de compuestas Chaptalia con especial referencia a las
especies argentinas. Darwiniana 6(4): 505594.
Hansen, H. V. (1990). Phylogenetic studies in the Gerbera-complex (Compositae, tribe Mutisieae,
subtribe Mutisiinae). Nordic J. Bot.
9(5): 469é--485.
Nesom, G. L. (1984). Taxonomy and distribution of Chaptalia dentata and C. albicans (Asteraceae: Mutisieae). Brittonia 36(4): 396401.
Nesom, G. L. (1995). Revision of Chaptalia (Asteraceae: Mutisieae) from North America and continental Central America. Phytologia 78(3): 153188.
Key to species
1. Leaves long-petiolate, petiole as long as or much longer than lamina Ch. rotundifolia
Leaves short-petiolate or sessile 2
2. (1) Leaves
lyrate or lyrate-pinntifid 3
Leaves entire or dentate 4
3. (2) Stipe ebracteolate; capitula 100300-flowered; rostrum much longer than achene body; plants relatively large and robust, to 60 cm tall Ch. nutans
Stipe bracteolate; capitula 2540-flowered; rostrum shorter than achene body; plants relatively small, 1020 cm tall Ch. similis
4. (2) Leaves usually entire, sometimes denticulate; stipe nodding when young, ebracteolate; florets 100200 Ch. integerrima
Leaf margins retrorsely serrate; stipe always erect, bracteolate; florets 20605
5. (4) Secondary veins prominent on underside of leaf; lamina papyraceous; achene body papillose Ch. mandonii
Secondary
veins almost invisible; lamina membranaceous; achene body glabrous or papillose
at base Ch. runcinata
Chaptalia
cordata Hieron. var. ferrugineo-tomentosa
Hieron., Bot. Jahrb. Syst. 36(5): 512 (1905) = *?Chaptalia rotundifolia D. Don
Chaptalia
diversifolia Greene, Leafl. Bot. Observ. Crit. 1: 194 (1906) = Chaptalia nutans (L.) Polak.
*Chaptalia ebracteata (Kuntze) K. Schum.,
Bot. Jarhb. Syst. 26(1): 376 (1900) = Chaptalia
nutans (L.) Polak.
Chaptalia
erosa Greene, Leafl. Bot. Observ. Crit. 1: 196 (1906) = Chaptalia nutans (L.) Polak.
Chaptalia
fibrosa Beauverd, Bull. Soc. Bot. Genve 2, ser. 13: 11 (1921), nom. nud.
(based on Herzog 1257) = untreated by
Burkart but determined as Chaptalia
piloselloides var. graminifolia
Malmeč'' (vide Koster, 1945: 675). The taxon cited Koster was treated as a
variety of C. runcinata by Burkart,
but considered endemic to Brazil.
Chaptalia graminifolia (Dusn) Cabrera, Fl. Ilust. Catarinense, Compostas Tribo 1. Mutisieae : 69 (1973) = Chaptalia runcinata Kunth
*Chaptalia
integrifolia (Cass.) Baker in Mart., Fl. Bras. 6(3): 377 (1884), comb.
illeg., non Chaptalia integrifolia
(Michx.) Nutt. (= Chaptalia tomentosa Vent.) = Chaptalia integerrima (Vell.) Burkart
Chaptalia integerrima
(Vell.) Burkart, Darwiniana 6(4): 576 (1944).
Tussilago integerrima
Vell., Fl. Flum. Icones 8, tab. 140 (1831). Type: not cited; the original of Fl.
Flum. Icones 8: tab. 140 (1831) is best treated as the lectotype in the event
that no material of Flora Fluminensis
is found.
Leria integrifolia
Cass., Dict. Sci. Nat. 26: 103 (1823). Types: Uruguay é''Nous avons fait cette
description sur deux chantillons secs de lherbier de M. de Jussieu,
recueillis par Commerson dans les
environs de Montevideo; lé''un de ces deux chantillons porte des fleurs, et
lautre des fruits m rs: ce dernier, tant plus grand que lautre dans toutes
ses parties, nous fait pr sumer que la plante acquiert de laccroissement aprs
la fleuraison. Type material: P-JU.
Leria nutans (L.) DC. var. integrifolia (Cass.) Less., Linnaea 5: 354 (1830).
Leria lutescens
Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865), nom. nud. (based on Mandon 10).
Leria lutescens
Sch.Bip., Linnaea 34(5): 527 (Feb. 1866). Type: č''[Mandon] 10. Leria lutescens Sz. Bip. a sp. aff. L. integrifoliae
Cass. sed capitula duplo minora. [Enumerato Cassiniacearum a cl. G. Mandon in Bolivia a. 18571861
lectarum]. Holotype P; isotypes: B, K, LE.
*Chaptalia integrifolia
(Cass.) Baker in Mart., Fl. Bras. 6(3): 377 (1884), comb. illeg., non Chaptalia integrifolia (Michx.) Nutt. (=
Chaptalia tomentosa Vent.)
Thyrsanthema
integrifolia (Cass.,) Kuntze, Revis. Gen. Pl. 3(3): 182 (1898).
*Chaptalia microdonta
Greene, Leafl. Bot. Observ. Crit. 1: 196 (1906). Type: Yungas, Bolivia, 1894, Miguel Bang, n. 2095. (Holotype US;
isotypes: B, K, US, Z).
Brazil: Bahia, Gois, Minas Gerais, Rio de Janeiro, S o
Paulo.
Argentina, Bolivia (La Paz, Santa Cruz), ?Guyana,
Paraguay, Peru, Uruguay.
Burkart (1944: 580) cited the following: Santa Cruz,
prov. Sara, Steinbach 6497 (BA, LIL);
Buena Vista, Steinbach 1574 (LIL).
Lengua de vaca;
Pelosilla (Burkart, 1944; Katinas,
1996); Lngua-de-vaca (Cabrera
& Klein, 1973); Vakak , Yty ar
(Cabrera, 1998); Lengua de vaca,
Pelosilla, Vakaku, Yerba de
*Chaptalia
integrifolia (Cass.) Baker in Mart., Fl. Bras. 6(3): 377 (1884), comb.
illeg., non Chaptalia integrifolia
(Michx.) Nutt. (= Chaptalia tomentosa Vent.) = Chaptalia integerrima (Vell.) Burkart
*Chaptalia
majuscula Greene, Leafl. Bot. Observ. Crit. 1: 196 (1906) = Chaptalia nutans (L.) Polak.
*Chaptalia
mandonii Burkart, Darwiniana 6(4): 551 (1944). Types: [Bolivia:] Larecaja,
en la vecind. de Sorata, monte Ch[?]heca a 3000 m, Mandon 11; X-1858, sub Leria
mandoni Schultz-Bip. (B., GOET., LE) (Typus).é'' Syntypes: B, GOET, K, LE,
P, S.
Leria mandoni
Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea, 34: 527 (Feb. 1866), nom.
nud. (based on Mandon 11)
*Chaptalia
piloselloides sensu Baker in Mart., Fl. Bras. 6(3): 378 (1884), p.p.
Brazil: Minas Gerais, So Paulo.
Argentina, Bolivia (La Paz), Brazil, Uruguay.
*Chaptalia
microdonta Greene, Leafl. Bot. Observ. Crit. 1: 196 (1906) = Chaptalia integerrima (Vell.) Burkart
*Chaptalia
nutans (L.) Hemsley, Biol. Centr. Amer. Bot. 2: 255 (1881) = Chaptalia nutans (L.) Polak.
Chaptalia nutans
(L.) Polak., Linnaea 41: 582 (1877).
Tussilago nutans
L., Syst. Nat. ed 10, 2: 1214 (1759). Type: Habitat in Americaă''. . Lectotype
(selected by Simpson in Woodson & Schery, 1975: 1278): Browne, Herb. Linn.
No. 995.5 (LINN). Note: Additional commentary has been provided by Nesom (1995:
162).
Tussilago lyrata
Pers., Syn. Pl. 2: 456 (1807). Type: not cited Hab. in Amer. meridionali..
The following specimens were cited by Cassini (1823: 103), les uns dans lile
de Saint-Domingue par M. Poiteau, les
autres dans lile de Porto-Rico par M.
Riedl. when the same species was transferred to Leria.
Leria nutans (L.)
Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 4 (1818).
Leria lyrata
(Pers.) Cass., Dict. Sci. Nat. 26: 102 (1823).
Tussilago vaccina
Vell., Fl. Flum. Icones 8: tab. 143 (1831). Type: not cited. [Fl. Flum. Ic. 8:
tab. 143].
Gerbera
nutans (L.) Sch.Bip. in Seem., Bot. Voy. Herald : 313 (1856).
*Chaptalia nutans (L.)
Hemsley, Biol. Centr. Amer. Bot. 2: 255 (1881), comb. illegit. superfl.
Thyrsanthema nutans
(L.) Kuntze, Revis. Gen. Pl. 1: 369 (1891).
Thyrsanthema ebracteata
Kuntze, Revis. Gen. Pl. 3(3): 182 (1898). Type: é''Bolivia: 2600 m Copachuncho in
Sierra de Santa Cruz. [Bolivia.
Copachuncho in Sierra de Santa Cruz, 2600 m, May 1892, Kuntze s.n. according to Wetter & Zanoni, 1985: 338].
Holotype: NY; isotype: B.
*Chaptalia ebracteata
(Kuntze) Schumann, Bot. Jahrb. Syst. 26: 376 (1898).
Chaptalia texana Greene,
Leafl. Bot. observ. Crit. 1: 191 (1906). Type: Rocky sparsley wooded ground in
western Texas, the type Neallys 297
as in U.S. Herb., Lindheimers n. 446
and the n. 674 of the Mexican Boundary Survey appear to be the same; perhaps
also Reverchons n. 1546, but that is
doubtful. It seems different. Holotype: US; isotype: SI.
Chaptalia
diversifolia Greene, Leafl. Bot. Observ. Crit. 1: 194 (1906).
Type: Near Mazarenango, Guatemala, 20 Febr., 1905 W.R. Maxon & R. Hay, n. 3504. Holotype US.
Chaptalia subcordata Greene,
Leafl. Bot. Observ. Crit. 1: 195 (1906). Type: St. Croix. Rickseckers n. 447. Holotype US.
Chaptalia
erosa Greene, Leafl. Bot. Observ. Crit. 1: 196 (1906). Type:
Costa Rica, June, 1892, Conduz
[sic!], n. 447 [sic!]. Holotype: US. (Nesom (1995) gave this as COSTA RICA.
San Jos, bords des chemins et fosses, 115 m, Jun 1892. A. Tonduz 4147.).
*Chaptalia
majuscula Greene, Leafl. Bot. Observ. Crit. 1: 196 (1906).
Types: é''Bolivian species. Rusbys n.
1677 from Mapiri and Bangs 237 from Yungas
.... Syntypes: US; isosyntype (Bang
237), K.
Chaptalia nutans
(L.) Polak. var. texana (Greene)
Burkart, Darwiniana 6(4): 569 (1944).
Argentina, Bolivia (La Paz, Santa Cruz), Brazil,
Colombia, Costa Rica, Dominican Republic, Ecuador, Guatemala, Guyana, Honduras,
Jamaica, Mexico, Panama, Paraguay, Peru, Puerto Rico, Uruguay, Venezuela.
Lngua-de-vaca, Costa-branca (Cabrera & Klein, 1973); Tapyi rapeo (Cabrera, 1998); Arracachuela, Burro-ca , Cerraja, Costa branca, Lechuguilla, Lengua de vaca, Peludilla, Pelusa, Raz de lombriz, Tapyi rapeo (Freire et al., 2006); Cerraja; Pelusa (Katinas, 1996).
Note: The combination Leria
nutans (L.) DC., Ann. Mus. Natl. Hist. Nat. 19: 68 (1812)í'' is cited in Index Kewensis and on the IPNI database
but the combination was never actually made by de Candolle, simply a comment
that several species of Tussilago
belonged in Leria!
*Chaptalia piloselloides (Vahl) Baker in Mart., Fl. Bras. 6(3): 378 (1884), p.p. = Chaptalia mandonii (Sch.Bip.) Burkart
Chaptalia
piloselloides (Vahl) Baker var. graminifolia
Dusn ex Malme, (1933) = Chaptalia
runcinata Kunth
*Chaptalia
piloselloides (Vahl) Baker in Mart., Fl. Bras. 6(3): 378 (1884), p.p. = Chaptalia runcinata Kunth
*Chaptalia
rotundifolia D. Don, Trans. Linn. Soc. London 16(2): 242
(1830). Type: In stagnatis altis frigidis Peruviae ad Pillao. Ruiz et Pavon. K.é'' Holotype: originally in Aylmer Bourke
Lamberts herbarium now presumably in BM see Miller (1970: 538540). Note:
the original Ruiz & Pavn collection is in MA, to be counted as an isotype
[sheet marked B8 of microfiche sheet 286 of the Ruiz & Pav n herbarium.
This sheet is labelled Perdicium lanatum/sp. n./ Pillao 1787 and on which are
mounted seven plants.]
Chaptalia cordata Hieron. var. ferrugineo-tomentosa Hieron., Bot. Jahrb. Syst. 36(5): 512 (1905). Type: Peruvia: crescit prope Cutervo (J.[elski] n. 726, m. Aprili 1879). Holotype: B.
Bolivia (?), Peru.
Note: The following synonymy is sometimes attributable to Chaptalia rotundifolia, but may be more appropriate under Trichocline T. ovalis is not present in Bolivia.
Chaptalia
ovalis D. Don, Trans. Linn. Soc. London 16(2): 241 (1830). Type:
In Peruvi ad Huassahuassi et Churapallanam. Ruiz et Pavon. K.
Holotype: originally in Aylmer Bourke LambertíYŤí˛'s herbarium now presumably in BM
see Miller (1970: 538540). Note: No material is immediately evident in the
microfiche of Ruiz & Pavn herbarium! It is possible that since Pav n sold
further material that the location of any duplicates of the type material needs
further investigation.
Perdicium ovale
Ruiz & Pav. ex D. Don, Trans. Linn. Soc. London 16(2): 241 (1830), nom.
nud. pro syn.
Trichocline ovalis
(D. Don) Hieron., Bot. Jahrb. Syst. 21(4): 370 (1896).
Chaptalia
runcinata Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed.
folio): 5 (1818). Type: [Colombia:] Crescit locis temperatis, scopulosis Andium Novo-Granatensium in ripa
fluvii Smita, alt. 590 hex. K Floret Octobri.. [Humboldt &
Bonpland 2031]. Holotype: P-Bonpl.; isotype: B-W.
Loxodon longipes Cass., Dict. Sci. Nat. 27: 255 (1823), nom.
illegit. pro Chaptalia runcinata
Kunth
Oxydon bicolor
Less, Linnaea 5: 357 (1830), nom. illegit. pro Chaptialia runcinata Kunth
*Chaptalia piloselloides (Vahl) Baker in Mart., Fl. Bras. 6(3): 378 (1884), p.p.
Chaptalia piloselloides (Vahl) Baker var. graminifolia Dusn ex Malme, Kongl. Svensk. Vetenskapsakad. Handl. 12(2): 115 (1933). Types: [Brazil:] [Paran ] Inter Capo Grande et Villa Velha 12/304 ([Dus n] n. 4062), ..., Curityba 2/909 ([Dusn] n. 8796), ..., Jaguariahyba 5/710 ([Dus n] 10037), 21/514 ([Dusn] n. 406a). Hab. in ripis rivulorum et campis plus minusve uliginosis. Note: In his discussion of this taxon Burkart (1944: 557) noted that a specimen in B was a duplicate of the type, this effectively indicating that Burkart lectotypified the name based on Dus n 4062; the lectotype is assumed to be in S. Burkart aslo cited a duplicated of Dusn 10037 in B, and duplicates of Dus n 406a in LE and SI.
Chaptalia runcinata Kunth var. graminifolia (Dusn ex Malme) Burkart, Darwiniana 6(4): 556 (1944).
Chaptalia graminifolia
(Dus n) Cabrera, Fl. Ilust. Catarinense, Compostas Tribo Mutisieae : 69 (1973).
Argentina, Bolivia (Santa Cruz), Brazil, Colombia, Costa Rica, Ecuador, Paraguay, Uruguay. Note: this species is not apparently recorded for Peru.
Lnguia-de-vaca,
L ngua-de-vaca-folha-de-grama (Cabrera & Klein, 1973); Cerraja (Cabrera, 1998).
Chaptalia similis
R. E. Fr., Nova Acta Regiae Soc. Sci. Upsal., ser. 4, 1: 95 (1905). Syntypes: [Argentina:]
Prov. Jujuy: S:a Catalina in fissuris rupium, ca. 3650 m. s. m. (1618 Jan.
1901; Kurtz 11460); Moreno in
fissuris rupium, 3500 m. s. m. (10 Dec. 1901; Fr[
Argentina, Bolivia (La Paz), Peru.
Note: Bukart (1944: 573) cited two collections from Bolivia,
noting that Mandon 13 (K) was a
cotype (= syntype), which it most certainly couldnt be since it was never
cited in the protologue!
Cited collections: Puna Patanca, Fiebrig 3197 (SI, LE, LIL, B); La Paz,
Potopoto, a 3700 m, Mandon 13; I-1861
(K).
Chaptalia
subcordata Greene, Leafl. Bot. Observ. Crit. 1: 195 (1906) = Chaptalia nutans (L.) Polak.
Chaptalia
texana Greene, Leafl. Bot. Observ. Crit. 1: 191 (1906) = Chaptalia nutans (L.) Polak.
Chatiakella
platyglossa Cass., Dict. Sci. Nat. 46: 403 (1827), based on Chylodia sarmentosa (Rich.) Rich. ex
Cass. = Tilesia baccata (L.f.)
Pruski
Chatiakella
stenoglossa Cass., Dict. Sci. Nat. 46: 403 (1827), based on Chylodia sarmentosa (Rich.) Rich. ex
Cass. = Tilesia baccata (L.f.)
Pruski
