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Belloa Remy, Fl. Chil. 3: 336 (1847).

Lucilia Cass. sect. Lucilioides DC., Prodr. 6: 46 (1838). Type: Lucilia chilensis Hook. & Arn. = Belloa chilensis (Hook. & Arn.) Remy

Merope Wedd., Chloris Andina 1: 160 (1856), p.p. nom. illegit., non Merope M. J. Roemer (1846) [RUTACEAE].

Lucilia Cass. sect. Lucilioides DC. subsect. Lucilioides S. E. Freire ser. Lucilioides S. E. Freire, Cladistics 3(3): 271 (1987).

Lucilia Cass. sect. Lucilioides DC. subsect. Lucilioides S. E. Freire ser. Paralucilia S. E. Freire., Darwiniana 27(14): 479 (1986). Type Lucilia pusilla (Kunth) Hieron. = Belloa kunthiana (DC.) Anderb. & S. E. Freire

Type: Belloa chilensis (Hook. & Arn.) Remy

Note: Anderberg & Freire (1991) proposed the continued recognition of Belloa and Luciliocline as separate from Lucilia. The diagnostic characters are rather weak but the genera are listed separately in this account.

References

Anderberg, A. A. & S. E. Freire. (1991). A cladistic and biogeographic analysis of the Lucilia group (Asteraceae, Gnaphalieae). Bot. J. Linn. Soc. 106(2): 173198.

Cabrera, A. L. (1958). El genero Belloa Remy. Bol. Soc. Argent. Bot. 7(2): 7985.

Dillon, M. O. & A. Sagstegui-Alva. (1991). Belloa. In: J. F. Macbride & collab., Flora of Peru, Family Asteraceae: Part V. Fieldiana, Bot. n.s. 26 (1422): 1118.

Freire, S. E. (1986). Revision del genero Lucilia (Compositae, Inuleae). Darwiniana 27(1--4): 431490.

Sagstegui-Alva, A. & M. O. Dillon. (1985). New species and combinations in Belloa (Inuleae-Asteraceae). Phytologia 58(6): 392400.

Key to species

1. Plants subcaespitose or erect; stems simple or with short branches; leaves -'lax or rosettiform on same plant or all approximate or all lax; inflorescences branched or in pseudoglomerules 2

Plants caespitose; stems simple, short; leaves rosettiform; capitula solitary in terminal rosette of leaves 4

2. (1) Stems branched; capitula in pseudoracemes B. schultzii

Stems simple; capitula in pseudospikes 3

3. (2) Outer phyllaries acuminate, inner phyllaries with fenestrate stereome; leaves pseudopetiolate B. pickeringii

Outer phyllaries acute; inner phyllaries with entire stereome; leaves not pseudopetiolate B. piptolepis

4. (1) Achenes with twin-hairs; marginal florets 3878; disc florets 48; leaves 618 mm long B. kunthiana

Achenes glabrous or with globose and mucilaginous hairs; marginal florets 125181; disc florets 1016; leaves 2050 mm long B. longifolia

Belloa argentea (Wedd.) Cabrerea, Fl. Prov. Jujuy 10: 296 (1978) = Belloa schultzii (Wedd.) Cabrera

Belloa burkartii (Cabrera) Cabrera, Revista Invest. Agr. 11: 404 (1957) = Luciliocline burkartii (Cabrera) Anderb. & S. E. Freire

Belloa caespititia (Wedd.) Cabrera, Bol. Soc. Argent. Bot. 7: 81 (1958) = Belloa schultzii (Wedd.) Cabrera

Belloa erythractis (Wedd.) Cabrera, Revista Invest. Agrc. 11: 404 (1957) = Gamochaeta erythractis (Wedd.) Cabrera

Belloa kunthiana (DC.) Anderb. & S. E. Freire, Bot. J. Linn. Soc. 106(2): 189 (1991).

Conyza pusilla Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 54 (1818), nom. illegit., non Houtt. (1773-1783). Type: [Ecuador:] Crescit regione alpina Regni Quitensis ? K'' Holotype: P-Bonpl.

Conyza kunthiana DC., Prodr. 5: 379 (1836). Type: [Ecuador:] K in regione alpin regni Quitensis? [Humboldt & Bonpland, s.n.] Holotype: P-Bonpl.

*Lucilia affinis Wedd., Chloris Andina 1: 230 (1857). Type: ''Hab. Bolivie!: Cordillres du d partement de la Paz (Mandon [168]). Holotype: P.

Lucilia conoidea Wedd., Chloris Andina 1: 154 (1856). Type: Hab. Prou!: Cordill res du dpartement de Cuzco (Gay). Holotype: P; isotype: LP.

*Lucilia violacea Wedd., Chloris Andina 1: 155 (1856). Type: ''Hab. Bolivie: sur les parties les plus lev es des montagnes de la Cordillres de Sorata!, la hauteur de 4000 5000 m tres (Wedd.). Holotype: P.

Merope kunthiana (DC.) Wedd., Chloris Andina 1: 161 (1856).

*Gnaphalium kunthianum (DC.) Kuntze, Revis. Gen. Pl. 3(3): 152 (1898).

Lucilia pusilla (Kunth) Hieron., Bot. Jahrb. Syst. 29(1): 29 (1900), comb. illegit.

Lucilia venezualensis Steyerm., Fieldiana, Bot. 28(3): 644 (1953). Type: [Venezuela: ] Type in herb. Chi. Nat. Hist. Mus., collected in dry pramo along upper headwaters of R o Tormero, below El Aguila and above Chachapo, state of Mrida, alt. 3650-3965 m., March 13, 1944, Julian A. Steyermark 55657, in dense clumps."" Holotype: F; isotypes MO, NY.

Lucilia kunthiana (DC.) Zardini, Ann. Missouri Bot. Gard. 74(2): 431 (1987).

Argentina, Bolivia (La Paz), Colombia, Ecuador, Peru, Venezuela.

Belloa longifolia (Cuatrec. & Aristeg.) Sagsteg. & Dillon, Phytologia 58(6): 396 (1985).

Lucilia longifolia Cuatrec. & Aristeg., Fl. Venez. : 367 (1964). Type: [VENEZUELA] camino a Pico Bol var, 15 km. al sudeste de Mrida, cerca del Refugio Moya, alt. 4.1004.300 m. Edo. M rida, Venezuela ) E. L. Little, Jr. 15725, Oct. 1953). Holotype: VEN.

?Bolivia (?), Colombia, Peru, Venezuela.

Belloa lopezmirandae Cabrera, Bol. Soc. Argent. Bot. 7: 83 (1958) = Luciliocline lopezmirandae (Cabrera) Anderb. & S. E. Freire

Belloa pickeringii (A. Gray) Sagsteg. & Dillon, Phytologia 58(6): 396 (1985).

Lucilia pickeringii A. Gray, Proc. Amer. Acad. Arts 5: 138 (1861). Type: High Andes of Peru'' [Collected on the United States South Pacific Exploring Expedition under Captain Wilkes]. Note: no distinction was made between this citation and that of the following variety.

Lucilia (Merope) pickeringii A. Gray var. b? minor A. Gray, Proc. Amer. Acad. Arts 5: 138 (1861). Type: see under synonym above.

Argentina, Bolivia (La Paz), Peru, Venezuela.

Belloa piptolepis (Wedd.) Cabrera, Bol. Soc. Argent. Bot. 7: 81 (1958).

Merope piptolepis Wedd., Chloris Andina 1: 162 (1856). Types: Hab. Prou: collines pierreuses, pr s de Maravillas!, dans le dpartement de Puno, h. 3900 m tres (Wedd. [4514]). Bolivie: dpartement de Potosi!, aux environs des lagunas (dOrbigny, n. 1371).'' Lectotype (selected by Freire, 1986: 472): Weddell 4514, P; isolectotype: LP. Syntype: dOrbigny 1371, GH.

Lucilia piptolepis Wedd., Chloria Andina : tab. 26B (1856), based on Merope piptolepis Wedd., Chloris Andina 1: 162 (1856).

Gnaphalium evacoides Sch.Bip. ex Klatt, Linnaea 42(1): 142 (1878). Type: Crescit prope Parama de Mucuchier, Columbia, leg. Moritz 1047. Holotype: B. Note: Solbrig (1965) also suggested material may well be in GH.

Gnaphalium piptolepis (Wedd.) Griseb., Abh. Knigl. Gess. Wiss. G ttingen 24(1): 186 (March-April 1879); Symb. Fl. Argent.: 186 (1879).

Argentina, Bolivia (Cochabamba, La Paz), Chile, Ecuador, Peru, Venezuela.

Belloa punae (Cabrera) Cabrera, Revista Invest. Agric. 11(4): 404 (1958) = Luciliocline subspicata (Wedd.) Anderb. & S. E. Freire

Belloa schultzii (Wedd.) Cabrera, Revista Invest. Agric. 11(4): 404 (1958)

Merope argentea Wedd., Chloris Andina 1: 163 (1856). Type: Hab. Bolivie: dparement de La Paz, dans les endroits pierreux et un peu humides de la partie sup rieure de la Quebrada de Chuquiaguillo! (Wedd.). Holotype: P.

Merope schultzii Wedd., Chloris Andina 1: 163 (1856). Type: Gnaphalium evacoides Schultz Bip., in Bonplandia, ann. 1856, p. 54. Hab. Prou: au sommet de la Cordill re dAyapata!, dans la province de Carayaba (Lechler, exsicc., n. 1984). Holotype: P; isotype: LP.

Merope virescens Wedd., Chloris Andina 1: 163 (1856). Type: Hab. Bolivie: dans les mmes lieux que le M. argentea! (Wedd.). Holotype: P; isotype: LP.

Merope caespititia Wedd., Chloris Andina 1: 164 (1856). Type: YT'Hab. Bolivie: sur les coteaux pierreux, autour des lagunas de Potosi! (dOrbigny, n. 1399). Holotype: P; isotypes: GH, LP.

*Lucilia schultzii (Wedd.) A. Gray, Proc. Amer. Acad. Arts 5: 138 (1862).

Lucilia schultzii Wedd. ex Sch.Bip., Linnaea, 34: 532 (Feb. 1866), nom. nud.

Gnaphalium schultzii (Wedd.) Cabrera, Not. Mus. La Plata, Bot., 13: 14 (1948), comb. illegit. non Mendonca (1943).

Gnaphalium viridescens Cabrera, Notas. Mus. La Plata, secc. Bot. 13(56): 15 (1948), non Gnaphalium virescens Kuntze

Belloa virescens (Wedd.) Cabrera, Revista Invest. Agric. 11(4): 404 (1958).

Belloa caespititia (Wedd.) Cabrera, Bol. Soc. Argent. Bot. 7: 81 (1958).

Belloa argentea (Wedd.) Cabrerea, Fl. Prov. Jujuy 10: 296 (1978).

Argentina, Bolivia (La Paz, Oruro), Chile, Peru.

Belloa subspicata Wedd., Chloris Andina 1: 159 (1856) = Luciliocline subspicata (Wedd.) Anderb. & S. E. Freire

Belloa virescens (Wedd.) Cabrera, Revista Invest. Agric. 11(4): 404 (1958) = Belloa schultzii (Wedd.) Cabrera

Bichenia auriculata Wedd., Chloris Andina 1: 26 (1855) = Trichocline auriculata (Wedd.) Hieron.

Bichenia reptans Wedd., Chloris Andina 1: 25, tab. 8, B (1855) = Trichocline reptans (Wedd.) B. L. Rob.