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Belloa Remy,
Fl. Chil. 3: 336 (1847).
Lucilia Cass. sect.
Lucilioides DC., Prodr. 6: 46 (1838).
Type: Lucilia chilensis Hook. &
Arn. = Belloa chilensis (Hook. &
Arn.) Remy
Merope
Wedd., Chloris Andina 1: 160 (1856), p.p. nom. illegit., non Merope M. J. Roemer (1846) [RUTACEAE].
Lucilia
Cass. sect. Lucilioides DC. subsect. Lucilioides S. E. Freire ser. Lucilioides S. E. Freire, Cladistics
3(3): 271 (1987).
Lucilia
Cass. sect. Lucilioides DC. subsect. Lucilioides S. E. Freire ser. Paralucilia S. E. Freire., Darwiniana
27(14): 479 (1986). Type Lucilia pusilla
(Kunth) Hieron. = Belloa kunthiana
(DC.) Anderb. & S. E. Freire
Type: Belloa chilensis (Hook. & Arn.) Remy
Note: Anderberg & Freire (1991) proposed the continued
recognition of Belloa and Luciliocline as separate from Lucilia. The diagnostic characters are
rather weak but the genera are listed separately in this account.
References
Anderberg, A. A. & S. E. Freire. (1991). A cladistic and biogeographic analysis of the Lucilia group (Asteraceae, Gnaphalieae). Bot. J. Linn. Soc. 106(2): 173198.
Cabrera, A. L. (1958). El genero Belloa Remy. Bol. Soc. Argent. Bot. 7(2): 7985.
Dillon, M. O. & A. Sagstegui-Alva. (1991).
Belloa. In: J. F. Macbride &
collab., Flora of Peru, Family Asteraceae: Part V. Fieldiana, Bot. n.s. 26
(1422): 1118.
Freire, S. E. (1986). Revision del genero Lucilia (Compositae, Inuleae). Darwiniana 27(1á--4): 431490.
Sagstegui-Alva, A. & M. O. Dillon.
(1985). New species and combinations in Belloa
(Inuleae-Asteraceae). Phytologia 58(6): 392400.
Key to species
1. Plants
subcaespitose or erect; stems simple or with short branches; leaves á-'lax or
rosettiform on same plant or all approximate or all lax; inflorescences
branched or in pseudoglomerules 2
Plants
caespitose; stems simple, short; leaves rosettiform; capitula solitary in
terminal rosette of leaves 4
2. (1) Stems
branched; capitula in pseudoracemes B. schultzii
Stems
simple; capitula in pseudospikes 3
3. (2) Outer
phyllaries acuminate, inner phyllaries with fenestrate stereome; leaves
pseudopetiolate B. pickeringii
Outer
phyllaries acute; inner phyllaries with entire stereome; leaves not
pseudopetiolate B. piptolepis
4. (1) Achenes
with twin-hairs; marginal florets 3878; disc florets 48; leaves 618 mm long B. kunthiana
Achenes
glabrous or with globose and mucilaginous hairs; marginal florets 125181; disc
florets 1016; leaves 2050 mm long B. longifolia
Belloa argentea (Wedd.) Cabrerea, Fl. Prov. Jujuy 10: 296 (1978) = Belloa schultzii (Wedd.) Cabrera
Belloa
burkartii (Cabrera) Cabrera, Revista Invest. Agr. 11: 404 (1957) = Luciliocline burkartii (Cabrera)
Anderb. & S. E. Freire
Belloa caespititia (Wedd.) Cabrera, Bol. Soc. Argent. Bot. 7: 81 (1958) = Belloa schultzii (Wedd.) Cabrera
Belloa erythractis (Wedd.) Cabrera, Revista Invest. Agrc. 11: 404 (1957) = Gamochaeta erythractis (Wedd.) Cabrera
Belloa kunthiana
(DC.) Anderb. & S. E. Freire, Bot. J. Linn. Soc. 106(2): 189 (1991).
Conyza pusilla Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 54 (1818), nom. illegit., non Houtt. (1773-1783). Type: [Ecuador:] Crescit regione alpina Regni Quitensis ? Kí'' Holotype: P-Bonpl.
Conyza kunthiana
DC., Prodr. 5: 379 (1836). Type: [Ecuador:] K in regione alpin regni
Quitensis? [Humboldt & Bonpland,
s.n.] Holotype: P-Bonpl.
*Lucilia affinis Wedd., Chloris Andina 1: 230 (1857). Type: â''Hab. Bolivie!: Cordillres du d partement de la Paz (Mandon [168]). Holotype: P.
Lucilia conoidea Wedd., Chloris Andina 1: 154 (1856). Type: Hab. Prou!: Cordill res du dpartement de Cuzco (Gay). Holotype: P; isotype: LP.
*Lucilia violacea Wedd., Chloris Andina 1: 155 (1856). Type: é''Hab. Bolivie: sur les parties les plus lev es des montagnes de la Cordillres de Sorata!, la hauteur de 4000 5000 m tres (Wedd.). Holotype: P.
Merope kunthiana (DC.) Wedd., Chloris Andina 1: 161 (1856).
*Gnaphalium kunthianum (DC.) Kuntze, Revis. Gen. Pl. 3(3): 152 (1898).
Lucilia pusilla (Kunth) Hieron., Bot. Jahrb. Syst. 29(1): 29 (1900), comb. illegit.
Lucilia
venezualensis Steyerm., Fieldiana, Bot. 28(3): 644 (1953). Type:
[Venezuela: ] Type in herb. Chi. Nat. Hist. Mus., collected in dry pramo
along upper headwaters of R o Tormero, below El Aguila and above Chachapo,
state of Mrida, alt. 3650-3965 m.,
Lucilia kunthiana (DC.) Zardini, Ann. Missouri Bot. Gard. 74(2): 431 (1987).
Argentina, Bolivia (La Paz), Colombia, Ecuador, Peru, Venezuela.
Belloa
longifolia (Cuatrec. & Aristeg.) Sagsteg. & Dillon, Phytologia
58(6): 396 (1985).
Lucilia
longifolia Cuatrec. & Aristeg., Fl. Venez. : 367 (1964). Type:
[VENEZUELA] camino a Pico Bol var, 15 km. al sudeste de Mrida, cerca del
Refugio Moya, alt. 4.1004.300 m. Edo. M rida, Venezuela ) E. L. Little, Jr. 15725, Oct. 1953). Holotype: VEN.
?Bolivia (?), Colombia, Peru, Venezuela.
Belloa
lopezmirandae Cabrera, Bol. Soc. Argent. Bot. 7: 83 (1958) = Luciliocline lopezmirandae (Cabrera)
Anderb. & S. E. Freire
Belloa pickeringii (A. Gray) Sagsteg. & Dillon, Phytologia 58(6): 396 (1985).
Lucilia pickeringii A. Gray, Proc. Amer. Acad. Arts 5: 138 (1861). Type: High Andes of Peruá'' [Collected on the United States South Pacific Exploring Expedition under Captain Wilkes]. Note: no distinction was made between this citation and that of the following variety.
Lucilia (Merope) pickeringii A. Gray var. b? minor A. Gray, Proc. Amer. Acad. Arts 5: 138 (1861). Type: see under synonym above.
Argentina, Bolivia (La Paz), Peru, Venezuela.
Belloa piptolepis (Wedd.) Cabrera, Bol. Soc. Argent. Bot. 7: 81 (1958).
Merope
piptolepis Wedd., Chloris Andina 1: 162 (1856). Types: Hab. Prou: collines pierreuses, pr s de
Maravillas!, dans le dpartement de Puno, h. 3900 m tres (Wedd. [4514]). Bolivie:
dpartement de Potosi!, aux environs des lagunas (dOrbigny, n. 1371).é'' Lectotype (selected by Freire, 1986: 472): Weddell 4514, P; isolectotype: LP.
Syntype: dOrbigny 1371, GH.
Lucilia piptolepis Wedd., Chloria Andina : tab. 26B (1856), based on Merope piptolepis Wedd., Chloris Andina 1: 162 (1856).
Gnaphalium
evacoides Sch.Bip. ex Klatt, Linnaea 42(1): 142 (1878). Type: Crescit
prope Parama de Mucuchier, Columbia, leg. Moritz
1047. Holotype: B. Note: Solbrig (1965) also suggested material may well be in
GH.
Gnaphalium piptolepis (Wedd.) Griseb., Abh. Knigl. Gess. Wiss. G ttingen 24(1): 186 (March-April 1879); Symb. Fl. Argent.: 186 (1879).
Argentina, Bolivia (Cochabamba, La Paz),
Chile, Ecuador, Peru, Venezuela.
Belloa
punae (Cabrera) Cabrera, Revista Invest. Agric. 11(4): 404 (1958) = Luciliocline subspicata (Wedd.) Anderb.
& S. E. Freire
Belloa
schultzii (Wedd.) Cabrera, Revista Invest. Agric. 11(4): 404 (1958)
Merope argentea Wedd., Chloris Andina 1: 163 (1856). Type: Hab. Bolivie: dparement de La Paz, dans les endroits pierreux et un peu humides de la partie sup rieure de la Quebrada de Chuquiaguillo! (Wedd.). Holotype: P.
Merope schultzii Wedd., Chloris Andina 1: 163 (1856). Type: Gnaphalium evacoides Schultz Bip., in Bonplandia, ann. 1856, p. 54. Hab. Prou: au sommet de la Cordill re dAyapata!, dans la province de Carayaba (Lechler, exsicc., n. 1984). Holotype: P; isotype: LP.
Merope
virescens Wedd., Chloris Andina 1: 163 (1856). Type: Hab. Bolivie: dans les mmes lieux que le M. argentea! (Wedd.). Holotype: P; isotype: LP.
Merope caespititia Wedd., Chloris Andina 1: 164 (1856). Type: íYTí²'Hab. Bolivie: sur les coteaux pierreux, autour des lagunas de Potosi! (dOrbigny, n. 1399). Holotype: P; isotypes: GH, LP.
*Lucilia
schultzii (Wedd.) A. Gray, Proc. Amer. Acad. Arts 5: 138 (1862).
Lucilia schultzii Wedd. ex Sch.Bip., Linnaea, 34: 532 (Feb. 1866), nom. nud.
Gnaphalium schultzii (Wedd.) Cabrera, Not. Mus. La Plata, Bot., 13: 14 (1948), comb. illegit. non Mendonca (1943).
Gnaphalium viridescens Cabrera, Notas. Mus. La Plata, secc. Bot. 13(56): 15 (1948), non Gnaphalium virescens Kuntze
Belloa virescens (Wedd.) Cabrera, Revista Invest. Agric. 11(4): 404 (1958).
Belloa caespititia (Wedd.) Cabrera, Bol. Soc. Argent. Bot. 7: 81 (1958).
Belloa
argentea (Wedd.) Cabrerea, Fl. Prov. Jujuy 10: 296 (1978).
Argentina, Bolivia (La Paz, Oruro), Chile,
Peru.
Belloa subspicata
Wedd., Chloris Andina 1: 159 (1856) = Luciliocline
subspicata (Wedd.) Anderb. & S. E. Freire
Belloa virescens (Wedd.) Cabrera, Revista Invest. Agric. 11(4): 404 (1958) = Belloa schultzii (Wedd.) Cabrera
Bichenia auriculata Wedd., Chloris Andina 1: 26 (1855) = Trichocline auriculata (Wedd.) Hieron.
Bichenia
reptans Wedd., Chloris Andina 1: 25, tab. 8, B (1855) = Trichocline reptans (Wedd.) B. L. Rob.
