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Barnadesia Mutis in L.f., Suppl. Pl. : 55 (1782).

Bacasia Ruiz & Pav., Fl. Peru. Chil. Prodr. : 105, t. 22 (1794). Type: Barnadesia spinosa L.f.

Xenophontia Vell., Fl. Flum.: 346 (1825)[7 Sept. - 28 Nov. 1829]. Type: Xenophontia caryophyl[l]a Vell. = Barnadesia caryophylla (Vell.) S. F. Blake var. caryophylla

Diacantha Less., Linnaea 5: 243 (1830). Barnadesia arborea Kunth = Barnadesia caryophylla (Vell.) S. F. Blake var. caryophylla

Barnadesia Mutis sect. Penthea D. Don, Trans. Linn. Soc. London 16(2): 280 (1830). Type: Barnadesia laxa D. Don = Barnadesia caryophylla (Vell.) S. F. Blake var. caryophylla

Penthea (D. Don) Spach, Hist. Veg. Phan. 10: 9 (1841).

Rhodactinea Gardner, London J. Bot. 6: 449 (1847). Barnadesia rosea Lindl. = Barnadesia caryophylla (Vell.) S. F. Blake var. caryophylla

Barnadesia Mutis subgen Penthea (D. Don) Chung., Rev. Barnadesia (Compositae-Mutisieae). : 36 (1965).

Type: Barnadesia spinosa L.f.

References

Chung, In-Cho (1965). Revision of Barnadesia (Compositae-Mutisieae). : 186.

Hind, D. J. N. (2001). A new species of Barnadesia (Compositae: Barnadesieae) from Bolivia. Kew. Bull. 56(3): 705710.

Urtubey, E. (1999). Revisin del g nero Barnadesia (Asteraceae: Barnadesioideae, Barnadesieae). Ann. Missouri Bot. Gard. 86(1): 57117.

Key to species (from Hind, 2001)

1. Leaves distinctly petiolate, alternate (never fasciculate); capitula clearly pedicellate, several in corymbs; disc florets with tubular corollas and plumose pappus (Barnadesia subgen. Bacasia) B. corymbosa

Leaves sessile or pseudopetiolate, rarely short-petiolate, obviously fasciculate, occasionally with some leaves alternate; capitula sessile or shortly pedicellate, solitary or few in corymbs; disc florets with ligulate or bilabiate corollas and pappus simple or barbellate (Barnadesia subg. Barnadesia) 2

2. (1) Marginal florets 13; disc florets usually 3 (sect. Penthea) 3

Marginal florets 8; disc florets usually 1 (sect. Pauciflora) 6

3. (2) Disc floret pappus shorter than or equal to corolla tube B. odorata

Disc floret pappus much longer than corolla tube 4

4. (3) Disc floret corollas bilabiate, occasionally ligulate; involucre turbinate; phyllaries 1012-seriate, shiny; spines 1228 mm, in pairs B. caryophylla

Disc floret corollas always ligulate; involucre campanulate; phyllaries 89-seriate, matt, moderately to densely pubescent; spines 319 mm when present, in pairs or fascicles 5

5. (4) Leaves always fasciculate, usually densely pubescent, often with long-ciliate margins; stem pubescence of long, straw-coloured hairs; phyllaries usually moderately short-pubescent with straw-coloured hairs; leaves 944 4ퟀ16 mm, elliptic or oblong, apices attenuate or obtuse, mucronate; spines 6.519 mm, in pairs or fascilces B. polyacantha

Leaves alternate or in fascicles of few (24) leaves, usually moderately pubescent, usually with short-ciliate margins; stem pubescence of short, greyish hairs; phyllaries moderately to densely long-pubescent, especially so towards apices of inner phyllaries; leaves 2575 10ז15 (20) mm, narrowly elliptic, apices long-acute to attenuate, never mucronate; spines to c. 3 mm, in pairs when present B. woodii

6. (2) Disc floret pappus shorter than or equal to corolla tube; basal anther appendages short-sagittate; disc floret pappus setae simple B. glomerata

Disc floret pappus much longer than corolla tube anther bases decurrent; disc floret pappus setae barbellate 7

7. (6) Disc floret corollas glabrous inside throat; capitula 2446 mm tall; phyllaries 79 (10)-seriate; disc floret corolla lobes glabrous; leaves (5)1014 mm wide B. pycnophylla

Disc floret corollas pubescent inside throat; capitula 4550 mm tall; phyllaries 1011-seriate; disc floret corolla lobes sparsely villous; leaves 36 mm wide B. macrocephala

Barnadesia berberoides Sch.Bip., Berberid. Amer. Austral. : 57 (2857), nom. nud. = Barnadesia pycnophylla Muschl.

Barnadesia caryophylla (Vell.) S. F. Blake, Proc. Biol. Soc. Wash. 38: 86 (1925).

var. caryophylla

Xenophontia caryophylla Vell., Fl. Flum.: 347 (1825)[7 Sept. - 28 Nov. 1829]. Type: [Brazil:] Habitat silvis, fruticetisque mediterraneis. Fl. Flum. Ic. 8: tab. 8 (1831). Neotype (selected by Chung, 1965: 39): BRAZIL: Goyaz: Gardner 4268, K; isoneotypes: BM, F, G, GH, NY, P, US.

Barnadesia laxa D. Don, Trans. Linn. Soc. London 16(2): 280 (1830). Type: In Brasili. Sello. Holotype: originally in Aylmer Bourke Lamberts⒒ Herbarium, location of this specimen now unknown BM, CGE, G, LE, or OXF fide Miller (1970: 541). Rich also purchased some Sellow material and this is now in Herb. Delessert, P.

Barnadesia rosea Lindl., Bot. Reg. 29 t. 29 (1843). Type: Cult. Duke of Northumberland ex South America. Herbarium specimen unknown.

Rhodactinea rosea (Lindl.) Gardner, London J. Bot. 6: 450 (1847).

Barnadesia dianthiflora Mart. ex Baker in Mart., Fl. Bras. 6(3): 365 (1884), nom. nud. pro syn

Barnadesia verpretorum Mart. ex Baker in Mart., Fl. Bras. 6(3): 365 (1884), nom. nud. pro syn.

Barnadesia rosea Lindl. var. macrospinosa Loefgr., Bol. Comm. Geog. So Paulo 12: 448 (1897). Type: [Brazil:] Herbario da Commiss o numero 346. . Habitam caapes e beira campos nos Estados de Goyaz, Minas e S. Paulo. Os exemplares da Commiss o foram colhidos em Mogy-Guass e Itapetininga. Holotype: SP.

*Barnadesia spinosa, sensu auct. non. L.f. Note: Foster (19050: 205) cited this name (non sensu auct.), which is most probably referable to B. caryophylla.

Brazil: Goi s, Mato Grosso, Minas Gerais, Rio de Janeiro, So Paulo.

Bolivia (La Paz), Peru.

Barnadesia corymbosa (Ruiz & Pav.) D. Don, Trans. Linn. Soc. London 16(2): 278 (1830).

Bacasia corymbosa Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. : 189 (1798). Type: Habitat in Andium nemoribus prope Mu a vicium. Floret Augusto, et Septembri. Lectotype (selected here): MA [specimen marked as B4 on sheet 285 of the microfiche of the Ruiz & Pavn herbarium labelled Genus novum./Bacasia corymbosa󖑒 with a subsidiary label marked 29/15 Herbarium Peruvianum and determined as Barnadesia corymbosa Don by Domke]; isolectotypes: MA [sheets marked as B3 & B5 on sheet 285 of the microfiche of the Ruiz & Pavn herbarium]. Note: Chung (1965) merely mentioned a type, having seen a duplicate in BM, and Ferreyra (1995) mentioned the holotype, without declaring which of the three sheets it corresponded to, and Ferreyra was followed by Urtubey (1999).

*Barnadesia venosa Rusby, Mem. Torrey Bot. Club 6(1): 69 (1896). Types: [Bolivia:] Espirito Santo, 1891 ([Bang] 1265). Also collected by Pearce at Mu a, 70008000 ft. Cited by Chung as Holotype: GH; isotypes: MO, US. apparently assuming the Bang collection was the only type.

Bolivia (Cochabamba, Santa Cruz), Peru.

Barnadesia dianthiflora Mart. ex Baker in Mart., Fl. Bras. 6(3): 365 (1884), nom. nud. pro syn = Barnadesia caryophylla (Vell.) S. F. Blake var. caryophylla

Barnadesia divaricata Griseb., Abh. Knigl. Ges. Wiss. G ttingen 24(1): 209 (March-April 1879); Symb. Fl. Argent. : 209 (1879) = Dasyphyllum brasiliense (Spreng.) Cabrera var. divaricatum (Griseb.) Cabrera

*Barnadesia glomerata Kuntze, Revis. Gen. Pl. 3(3) 135 (1898). Type: Bolivia: Tunarigebirge. [Bolivia. Tunarigebirge, 2600 m, Apr-May 1892, Kuntze s.n. (2 sheets). according to Wetter & Zanonia, 1985: 327]. Lectotype (selected by Chung, 1965: 33): NY.

var. glomerata

Bolivia (Cochabamba)

var. mucronata Chung, Rev. Barnadesia : 33 (1965). Type: BOLIVIA: Santa Cruz: Comarapa, Jungas de San Mateo, Steinbach 8400. Holotype: GH; isotypes: BM, F, K, NY, S, US.

Bolivia (Santa Cruz).

Barnadesia hirsuta Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea 34(5): 527 (Feb. 1866), nom. nud. = Barnadesia polyacantha Wedd. var. velutina Chung

*Barnadesia inermis Rusby, Bull. New York Bot. Gard. 4(14): 399 (1907) = Dasyphyllum inerme (Rusby) Cabrera

Barnadesia laxa D. Don, Trans. Linn. Soc. London 16(2): 280 (1830) = Barnadesia caryophylla (Vell.) S. F. Blake

*Barnadesia macrocephala Kuntze, Revis. Gen. Pl. 3(3): 135 (1898). Type: Bolivia: 3000 m Sierra vor Colomi zwischen Cochabamba und Rio Juntas. [Bolivia. Sierra vor Colomi zwischen Cochabamba und Rio Juntas, 3000 m, 14 Apr 1892, Kuntze s.n. according to Wetter & Zanoni, 1985: 327]. Lectotype (selected by Chung, 1965: 32): NY.

Bolivia (Cochabamba).

Barnadesia odorata Griseb., Abh. Knigl. Ges. Wiss. G ttingen 24(1): 210 (March-April 1879), Symb. Fl. Argent.: 210 (1879). Types: [Argentina:] T.: Tucuman passim. J.: frequens. Syntype: Lorentz & Hieronymus 1013, 8. 5. 1873, GOET. Lectotype (selected by Chung, 1965: 38): G. Note: Chung lectotype citation was written BOLIVIA: Tucuman: Tuscan and Taruca Pampa near La Cruz, Lorentz & Hieronymus Dec. 22-28, 1872., not noting any other syntype collections.

Argentina, Bolivia (Cochabamba, Santa Cruz, Tarija).

*Barnadesia polyacantha Wedd., Chloris Andina 1: 13 (1855). Types: Hab. Bolivie: Andes de Sorata!, une hauteur de 2600 3300 mtres, sur les deux versants (Wedd.); province de Yungas? (dOrbigny).ퟙ Lectotype (selected by Chung, 1965: 43): Weddell 4468, P; isolectotypes: F, GH.

Bolivia (Cochabamba, La Paz, Laracaja, Santa Cruz), Peru.

var. polyacantha

var. velutina Chung, Rev. Barnadeisa : 46 (1965). Type: BOLIVIA: Larecaja: near Ananea, alt. 2400 m., Mandon 5. Holotype: BM; isotypes: F, G, K, NY, P.

Barnadesia hirsuta Sch.Bip., Bull. Soc. Bot. France 12: 79 (1865); Linnaea 34(5): 527 (Feb. 1866), nom. nud. (based on Mandon 5).

Bolivia (Larecaja).

Barnadesia pycnophylla Muschl., Bot. Jahrb. Syst. 50(2/3), Beibl. 111: 101 (1913). Type: Peruvia: VIa inter Tambo et Apurimac flumen in Departimento Ayacucho, provincia Huanta, frutices parvi in formatione herbis et graminibus compositae, 35003600 m s.m. (Weberbauer n. 5558. Florens 31. Maii 1910.) Holotype: B. Neotype (selected by Chung, 1965: 29): PERU: 6 miles south of Mito, Macbride & Featherstone 1830., US (1186011).

Barnadesia berberoides Sch.Bip., Berberid. Amer. Austral. : 57 (1857), nom. nud.

Bolivia (Cochabamba, La Paz), Peru.

Barnadesia rosea Lindl., Bot. Reg. 29 t. 29 (1843) = Barnadesia caryophylla (Vell.) S. F. Blake

Barnadesia rosea Lindl. var. macrospinosa Loefgr., Bol. Comm. Geog. So Paulo 12: 448 (1897) = Barnadesia caryophylla (Vell.) S. F. Blake var. caryophylla

*Barnadesia seleriana Muschl., Bot. Jahrb. Syst. 50(2/3), Beibl. 111: 99 (1913) = Dasyphyllum ferox (Wedd.) Cabrera

*Barnadesia spinosa auct. non L.f. = Barnadesia caryophylla (Vell.) S. F. Blake

*Barnadesia venosa Rusby, Mem. Torrey Bot. Club 6(1): 69 (1896) = Barnadesia corymbosa (Ruiz & Pav.) D. Don

Barnadesia verpretorum Mart. ex Baker in Mart., Fl. Bras. 6(3): 365 (1884), nom. nud. pro syn. = Barnadesia caryophylla (Vell.) S. F. Blake var. caryophylla

Barnadesia woodii D. J. N. Hind, Kew Bull. 56(3): 705 (2001). Type: Bolivia: La Paz State: Saavedra. Rio Comata Valley, on descent from Charazani on road to Apollo, alt. 1600 m, 6 Aug. 2000, Wood & Wasshausen 16594㑒. Holotype: K; isotype: LPB.

Bolivia (La Paz).