III. Subfam. Ajugoideae Kostel. (1834).

Subfam. Teucrioideae (Dumort.) Caruel (1884).


Low shrubs and perennial herbs, sometimes prostrate, decumbent or scandent, slightly aromatic; stems sometimes winged; leaves opposite, petiolate, simple or occasionally 3-lobed, ovate to triangular or suborbicular, dentate to crenate, sometimes caducous; inflorescence of axillary cymes, 1-5 flowered; cleistogamous [cl.] flowers sometimes present, minute, with corolla and stamens shorter than calyx; calyx (in both cl. and chasmogamous [ch.] flowers) actinomorphic or occasionally with 1 lobe smaller than others, 4-5-lobed, lobes broadly ovate to linear-lanceolate; corolla of ch. flowers purple, pink or white, zygomorphic, unlipped or 1-lipped with anterior lip present, 5-lobed, 4 posterior lobes similar, anterior lobe larger, often (always?) concave, corolla in bud asymmetrical; stamens of ch. flowers 4, didynamous, exserted, filaments arched or ±straight, thecae parallel to divaricate, confluent at dehiscence; pollen with supratectal spinules, columellae branched or granular; ovary unlobed to very shallowly 4-lobed; stigma-lobes equal to subequal; disc poorly developed or absent; nutlets obovoid, reticulately ridged, pubescent, attachment-scar lateral, 0.5-0.9x as long as nutlet.  Five species of forests and thickets in China.


Native to:

36 China


Chienodoxa Y.Z.Sun, Acta Phytotax. Sin. 1: 22 (1951).


Schnabelia Hand.-Mazz., Kaiserl. Akad. Wiss. Wien, Math.-Naturwiss. Kl., Anz. 58: 92 (1921); Cantino, Wagstaff & Olmstead, Syst. Bot. 23: 369-386 (1999).


Image resource

© Royal Botanic Gardens, Kew