N.B. the name of the author should have an accent, however this has been removed as it corrupts the program used to create the factsheets. 

III. Subfam. Ajugoideae Kostel. (1834).

Subfam. Teucrioideae (Dumort.) Caruel (1884).


Rhizomatous perennial herb, normally unbranched; leaves opposite, long-petiolate, broadly ovate, trilobed or trifoliolate, coarsely crenate to dentate; inflorescence a terminal thyrse; calyx 2-lipped (3/2), broadly campanulate, gaping, accrescent, in fruit gibbous, lips shallowly lobed; corolla white, 2-lipped (2/3) or 1-lipped (0/5), anterior lobe largest, corolla in bud asymmetrical; stamens 4, didynamous, long-exserted, filaments arched, thecae divergent to divaricate, confluent at dehiscence; pollen with supratectal spinules, columellae branched or granular; ovary very shallowly 4-lobed to unlobed; stigma-lobes equal; disc absent; nutlets broadly obovoid to subglobose, smooth to faintly ridged, glabrous, attachment-scar lateral, 0.7-1.0x as long as nutlet; endosperm scanty.  Two species of forests and open slopes in China and India and Myanmar.


Native to:

36 China
38 Eastern Asia
40 Indian Subcontinent
41 Indo-China


Cardioteucris C.Y.Wu, Acta Bot. Sin. 10: 247 (1962).


Rubiteucris Kudo, Mem. Fac. Sci. Taihoku Imp. Univ. 2 (2): 297 (1929); Cantino, Wagstaff & Olmstead, Syst. Bot. 23: 369-386 (1999).


Image resource

Royal Botanic Gardens, Kew