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Subfamily

VII. Subfam. Nepetoideae (Dumort.) Luerss. (1882).

VII. 1. Tribe Elsholtzieae (Burnett) Sanders & Cantino (1984).

Description

Robust annual herbs, strongly aromatic; leaves toothed, long-petiolate, in cultivated forms often deep purple-tinged or laciniate; inflorescence raceme-like, lax or congested, secund, of 2(-4)-flowered verticillasters; bracts small, often broadly ovate-acuminate; bracteoles absent; flowers small, distinctly pedicellate; calyx campanulate, lobes triangular, posterior lip shorter with median lobe shortest, tube densely hairy outside, annulate within throat, calyx strongly accrescent in fruit, with base of tube strongly distended and gibbous, calyx-lobes becoming slightly upcurved; corolla small, white to purplish, weakly 2-lipped, lobes subequal, corolla-tube very short, included in calyx, annulate at insertion of stamens; stamens 4, subequal, divergent, shorter than or equalling corolla, thecae separate, parallel, becoming slightly divergent, connective distinct; style short; nutlets 4, subglobose, glabrous, distinctly reticulate,  2n = 20, 40.  One variable species: P. frutescens (L.) Britton, probably native in shady areas in the Himalayan foothills, from Pakistan eastwards, but cultivated and occurring as a casual in SE Asia to China and Japan.

Distribution

Native to:

31 Russian Far East
36 China
38 Eastern Asia
40 Indian Subcontinent
41 Indo-China
42 Malesia

Introduced to:

14 Eastern Europe
33 Caucasus

Synonyms

Dentidia Lour., Fl. Cochinch.: 869 (1790).

Publication

Perilla L., Gen. Pl. ed 6, 578 (1764); Li Hsi-Wen & Hedge, Lamiaceae, in Wu Zheng-yi and Raven (eds). Flora of China 17: 241-242 (1994).

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Image resource

Royal Botanic Gardens, Kew