VII. Subfam. Nepetoideae (Dumort.) Luerss. (1882).
VII. 3. Tribe Ocimeae Dumort. (1829).
VII. 3. C. Subtribe Hyptidinae Endlicher (1838).
Trees, shrubs or subshrubs, often aromatic, hairs often branched; leaves simple, toothed; inflorescence of terminal or axillary thyrsoid, bracteate panicles, or axillary cymes; cymes (1-)3-many-flowered; bracts often leaf-like, sometimes conspicuous with white or purplish hairs; bracteoles inconspicuous; flowers shortly pedicellate; calyx tubular to infundibuliform, straight, actinomorphic to weakly zygomorphic, 5-lobed, lobes equal or slightly unequal with posterior lobe longer, erect or spreading; corolla strongly 2-lipped, 5-lobed (2/3), bluish-purple to lilac or rarely white, anterior lip with median lobe elongate, lateral lobes spreading, corolla-tube cylindrical to infundibuliform ( Sect. Umbellaria), straight; stamens filaments hairy; style with stylopodium overtopping ovary; disc weakly 4-lobed, lobes usually alternating with ovary lobes, but opposite in H. asperrima (Spreng.) Harley (Rudall 1981); nutlets ovoid to slightly flattened, sometimes winged ( Sect. Hyptidendron), weakly mucilaginous. 2n = c. 64. About 19 species in savanna and margins of montane forest, mainly central, eastern and northeastern Brazil, extending to Bolivia, with H. arborea (Benth.) Harley occurring in the Guayana Highlands: Venezuela, Guyana and northern Brazil, and disjunctly in the mountains of Colombia (Antioquia) and the Peru/Bolivia border. Two sections have been recognized (Harley 1988b): Sect. Hyptidendron (five species) with corolla-tube cylindrical, 6-11 mm long, nutlets narrowly winged and Sect. Umbellaria (Benth.) Harley (c. 14 species), corolla-tube <7mm long, ± infundibular, nutlets unwinged. H. arborea ( Sect. Hyptidendron) can attain a height of 20 m, though usually less.
82 Northern South America83 Western South America84 Brazil
Siagonarrhen Mart. ex J.A.Schmidt in C.F.P.von Martius & auct. suc. (eds.), Fl. Bras. 8: 146 (1858).
Hyptidendron Harley, Bot. J. Linn. Soc. 98: 90 (1988), rev.
Hyptis Jacq. sect. Buddleioides Benth. (1833).
Hyptis Jacq. sect. Umbellaria Benth. (1833).
© Royal Botanic Gardens, Kew