GrassBase - The Online World Grass Flora

Descriptions

W.D. Clayton, M. Vorontsova, K.T. Harman & H. Williamson

© Copyright The Board of Trustees, Royal Botanic Gardens, Kew.

Guadua

HABIT Perennial. Rhizomes short (27), or elongated (1); pachymorph. Culms erect (18/26), or geniculately ascending (1/26), or arching (5/26), or leaning (2/26), or scandent (4/26); 300–1145–3000 cm long; woody; without nodal roots (1/20), or with aerial roots from the nodes (1/20), or with root thorns from the nodes (19/20). Culm-internodes terete (22/27), or semiterete (5/27). Culm-nodes without obvious supra-nodal ridge (2/7), or with distinct supra-nodal ridge (5/7). Lateral branches dendroid. Branch complement one (4/20), or two (8/20), or three (13/20), or several (10/20), or many (1/20); in a horizontal line (1/5), or in a clump (4/5); with 1 branch dominant (9/9); thinner than stem (11/11). Culm-sheaths persistent (2/12), or tardily deciduous (1/12), or deciduous but leaving a persistent girdle (1/12), or deciduous (9/12); without auricles (13/16), or auriculate (6/16). Culm-sheath blade inconspicuous (1/18), or lanceolate (4/18), or ovate (2/18), or triangular (12/18); as wide as sheath at base (2/2). Leaf-sheath auricles absent (21), or erect (2), or falcate (5). Ligule an eciliate membrane (21), or a ciliolate membrane (10). Leaf-blade base with a brief petiole-like connection to sheath. Leaf-blades persistent (25), or deciduous at the ligule (3); linear (9), or lanceolate (26), or elliptic (1), or oblong (3), or ovate (2). Leaf-blade midrib conspicuous (5/5). Leaf-blade venation without cross veins (20), or with obscure cross veins (1), or with distinct cross veins (8).

INFLORESCENCE Synflorescence simple (1/26), or bractiferous (25/26); scanty (9/25), or paniculate (1/25), or globose (1/25), or clustered at the nodes (15/25); in untidy tufts (14/14); with glumaceous subtending bracts (24/25), or spathaceous subtending bracts (1/25); with axillary buds at base of spikelet (20/20); leafy between clusters (5/15), or leafless between clusters (8/15), or leafless between branches (2/15). Inflorescence a panicle (1/1).

Spikelets appressed (3/4), or ascending (4/4), or spreading (1/4), or deflexed (1/4). Fertile spikelets sessile (26/26).

FERTILE SPIKELETS Spikelets comprising 1 basal sterile florets (5/5), or 2 basal sterile florets (4/5); 1–6–17 fertile florets; with a barren rhachilla extension (1/26), or with diminished florets at the apex (26/26). Spikelets linear (16/26), or lanceolate (10/26), or elliptic (1/26), or oblong (3/26); laterally compressed (19/26), or subterete (7/26); 3–38.49–170 mm long; breaking up at maturity (26/26); disarticulating below each fertile floret (26/26). Rhachilla internodes definite (25/25); glabrous (18/25), or pubescent (7/25).

GLUMES Glumes two (6/22), or several (16/22); persistent (21/22), or deciduous (1/22); shorter than spikelet (22/22). Lower glume ovate (7/7); 0.25–0.5167–0.8 length of upper glume; membranous (1/4), or coriaceous (3/4); without keels (3/3); 7–9 -veined (2/4), or 10 -veined (1/4), or 11 -veined (3/4), or 12–13 -veined (2/4), or 14–17 -veined (1/4). Lower glume surface smooth (21/22), or asperulous (1/22); without pits (22/22); glabrous (20/22), or pubescent (2/22). Lower glume apex obtuse (3/6), or acute (4/6), or rostrate (1/6); muticous (6/7), or mucronate (1/7). Upper glume lanceolate (2/17), or ovate (16/17); 0.5–0.65–0.8 length of adjacent fertile lemma; coriaceous (11/11); without keels (12/12); 7–9 -veined (2/9), or 10 -veined (1/9), or 11–12 -veined (6/9), or 13 -veined (7/9), or 14–15 -veined (6/9), or 16–17 -veined (5/9), or 18–19 -veined (1/9). Upper glume lateral veins with cross-veins (4/4). Upper glume surface smooth (21/22), or asperulous (1/22); glabrous (19/22), or puberulous (3/22), or pubescent (3/22). Upper glume apex obtuse (2/13), or acute (11/13), or rostrate (1/13); muticous (12/17), or mucronate (5/17).

FLORETS Basal sterile florets barren (5/5); with palea (4/5), or without significant palea (2/5). Lemma of lower sterile floret lanceolate (2/5), or elliptic (1/5), or ovate (3/5); chartaceous (2/2); 10–14 -veined (1/4), or 15–16 -veined (2/4), or 17–20 -veined (1/4), or 23–27 -veined (1/4); obtuse (1/4), or acuminate (3/4); mucronate (5/5). Fertile lemma lanceolate (5/25), or elliptic (1/25), or ovate (22/25); coriaceous (26/26); without keel (26/26); wingless (26/26); 7–17–30 -veined. Lemma lateral veins obscure (1/5), or prominent (4/5). Lemma surface smooth (25/26), or scaberulous (1/26); unwrinkled (26/26); without grooves (26/26); glabrous (17/26), or puberulous (6/26), or pubescent (6/26), or pilose (2/26). Lemma margins flat (25/26), or convolute (1/26); exposing palea (25/26), or covering most of palea (1/26); eciliate (20/26), or ciliolate (3/26), or ciliate (3/26). Lemma apex obtuse (2/25), or acute (17/25), or acuminate (3/25), or apiculate (3/25); muticous (14/26), or mucronate (12/26), or awned (2/26); 1 -awned (1/1). Palea 0.75–0.9729–1 length of lemma; 6 -veined (1/9), or 8 -veined (2/9), or 9 -veined (6/9), or 10–11 -veined (5/9), or 12 -veined (3/9), or 13 -veined (2/9). Palea keels winged (26/26); eciliate (16/26), or pubescent (1/26), or ciliolate (6/26), or ciliate (3/26). Palea surface glabrous (20/26), or puberulous (1/26), or pubescent (5/26). Apical sterile florets resembling fertile though underdeveloped (23/26), or distinct from fertile (3/26); 1 in number (3/3); barren (1/1); rudimentary (2/3), or linear (1/3).

FLOWER Lodicules 3 (26/26); membranous (3/3); glabrous (19/26), or ciliate (7/26). Anthers 3 (2/26), or 6 (24/26). Stigmas 3 (26/26). Ovary unappendaged (19/26), or umbonate (7/26); pubescent on apex (2/3), or pubescent all over (1/3).

FRUIT Caryopsis with adherent pericarp (25/26), or fleshy pericarp (1/26); fusiform (4/7), or oblong (2/7), or ovoid (1/7), or obovoid (1/7). Embryo 0.2–0.2875–0.5 length of caryopsis. Hilum linear (1/1); 1 length of caryopsis.

DISTRIBUTION North America (6), or South America (27).

Please cite this publication as detailed in How to Cite Version: 27th January 2014.